臺灣博碩士論文加值系統

台灣的山蛩目馬陸始於高桑良興與王友燮的研究，但都於平原採集，1964之後再無人研究。整裡前人的研究與Korsós 2004年的名錄與未發表的資料，台灣山蛩目記錄6科7屬9種，但過去有些特徵描述與繪圖未竟詳細，且大部分並無留存標本可供對照。本研究即針對此目，以現有採集之標本做一分類研究。馬陸的採集範圍包含台灣本島與離島，採集到的馬陸在野外拍照記錄體色之後以75%乙醇保存，同時亦檢視國內其他研究單位的標本。所有標本透過解剖顯微鏡檢視，著重於文獻所描述的形態特徵包括領節形狀、足基節突起、臭孔位置、體節外表的花紋和生殖肢進行比對。重要的特徵均以拍照及繪圖的方式呈現。根據比較的結果，發現山蛩屬Spirobolus內有1種新亞種，4個新種，分別為S. panmaus sp. nov., S. taimalius sp. nov., S. redpodus sp. nov., S. lienhuachihus sp. nov. 及 formosae semiflavus subsp. nov.；將一物種復原且改變屬名派定Leptogoniulus takahasii (Takakuwa, 1940) stat. rev., comb. nov.；而根據先前所記載的9種中有3種目前尚未採集到，分別為Salpidobolus oceanicus, Spirobolus walkeri 及Spirobolus bungii。其餘7種經過比對與文獻相符和，故目前台灣可鑑定出的山蛩目應有4科5屬12種。

The taxonmic study of Spirobolida in Taiwan can be traced to the works done by Takakuwa and Wang from the 1940s, with most specimens collected from the plain. After their works, no other studies were done for the Taiwan millipede ever since. According to Korsós (2004) and unpublished information, there are 10 species in 7 genera and 6 families of spirobolids from Taiwan. However descriptions and illustrations of these species are insufficient and specimens are lacking for comparison. This study of Spirobolida is based on specimens collected from Taiwan recently and adjacent islands recorded color photos, and allocated in museums. Diagnostic Morphological characters include collum, coxae of legs, position of ozopore and gonopod. Four new species and 1 new subspecies of Spirobolus were designated in this study, S. panmaus sp. nov., S. taimalius sp. nov., S. redpodus sp. nov., S. lienhuachihus sp. nov. and S. formosae semiflavus subsp. nov.; a species revived and reassigned to other genus: Leptogoniulus takahasii (Takakuwa, 1940) stat. rev., comb. nov.; and 3 species reported earlier were not found, Salpidobolus oceanicus, Spirobolus walkeri and Spirobolus bungii. So far, there are 12 species, 5 genera, 4 family of Spirobolida in Taiwan.

誌謝
摘要
Abstract
Index
Introduction…………………………………………………………………………..1
Material and Methods.…………………………………………………………….....4
Taxonomic Accounts………………….………………………………………..….....7
Family Trigoniulidae Attems, 1909.......................................................................8
Genus Leptogoniulus Silvestri, 1897.............................................................9
Leptogoniulus sorornus (Butler, 1897)................................................10
Leptogoniulus takahasii (Takakuwa, 1940) comb. nov. .....................14
Genus Trigoniulus Pocock, 1894.................................................................17
Trigoniulus corallinus (Gervais, 1842)................................................18
Trigoniulus tertius Takakuwa, 1940....................................................21
Family Spirobolellidae Brölemann, 1913............................................................24
Genus Paraspirobolus Brölemann, 1902.....................................................26
Paraspirobolus lucifugus (Gervais, 1836)...........................................27
Family Pseudospirobolellidae Brölemann, 1913.................................................30
Genus Pseudospirobolellus Carl, 1912........................................................31
Pseudospirobolellus avernus (Butler, 1872)........................................32
Family Spirobolidae Bollman, 1893....................................................................35
Genus Spirobolus Brandt, 1833...................................................................37
Spirobolus formosae formosae Keeton, 1960......................................39
Spirobolus formosae semiflavus subsp. nov. ......................................41
Spirobolus panmaus sp. nov. ..............................................................43
Spirobolus taimalius sp. nov. ..............................................................46
Spirobolus redpodus sp. nov. ..............................................................48
Spirobolus lienhuachihus sp. nov. ......................................................50
Literature Cited.......................................................................53
Figures........................................................................56

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