本論文主要從事黃緣螢（Luciola ficta Olivier, 1909）幼蟲野放試驗及室內幼蟲之食性探討。以台北市信義區虎山溪作為試驗野放黃緣螢的地點，期望從復育黃緣螢的過程中，除了藉由清溪、教育宣導、棲地營造和釋放黃緣螢幼蟲（以下簡稱幼蟲）的活動中，促進社區民眾對生態環境的瞭解和認同，提高大眾保護自然環境的意願，同時進行螢火蟲的野放生態研究，累積棲地管理上的實務經驗。經過底質、流速和溪流動物的調查評估之後，選擇在主流旁的鋪石平台上小山溝作為主要野放區域，於1998年2月22日野放700隻3-5齡幼蟲；野放後每個星期進行幼蟲數量的追蹤。在4月中旬以標識再捕法估計成蟲數量，一夜最高的成蟲數量估計達44隻，並發現雌蟲在蘚苔植物上產卵，其後發現到有第二代幼蟲，但直到論文截稿為止，未發現第三代的幼蟲。進行野放過程出現的許多有關棲地管理的問題，例如：各級政府及基層民眾間之溝通、水質污染、棲地破壞及燈光管制等。在室內試驗部份，除了黃緣螢幼蟲各齡頭殼寬增長比例符合戴爾法則（Dyar''s rule）以外，頭殼長度和前胸背板長寬度均呈幾何級數的增長，亦符合戴爾法則（Dyar''s rule），幼蟲各齡外部形態的差異以及頭殼和前胸背板的增長特性可作為判斷齡期的依據。在25℃光照條件12D：12L生長箱中，黃緣螢幼蟲共有6個齡期，1~ 6齡幼蟲發育時間各為26.1 ± 0.6、25.0 ± 1.1、36.3 ± 1.2、45.7 ± 2.6、61.3 ± 2.5和52.4 ± 6.9天，共270.5天。在食物偏好試驗的結果，發現在所供應的三種食物（拉都希蛙的蝌蚪、褐吻蝦虎、石田螺）中，南港幼蟲對蝌蚪有取食偏好；陽明山和南港的幼蟲對所供應的六種螺肉（囊螺、石田螺、中國福田螺、福壽螺、川蜷、臺灣蜆）無取食偏好；六齡幼蟲取食殼寬在1.6 ~ 2.4 mm，殼高2.8 ~ 10.9 mm之間的川蜷；而從五種螺類的試驗中，發現幼蟲對螺的種類並無顯著的取食偏好，推測幼蟲捕食螺類的因子在於螺類是否容易捕獲，而非螺的種類。

Study on ecology of aquatic firefly, Luciola ficta Olivier, were carried out in the field and the laboratory. The fireflies were reintroduced in the Hu-Shan Creek, Taipei by means of reconstructing their habitat and releasing their larvae to establish its population and restore its habitat. Seven hundred larvae of the 3rd ~ 5th instars were released in a runnel aside the Creek, on Feb. 22th, 1998, and the numbers of larvae were monitored weekly. The numbers of adults were estimated daily by mark-recaptured method from April 16th to April 23th. The estimated number of adults reached maximum (44) on April 18th. The females were observed to lay eggs on the bryophytes. The laid eggs hatched, and these larvae perhaps grew up to adults in the field, but didn''t last to the next generation. Water pollution, habitat destruction, and streetlight were regarded as the major reasons that the fireflies were unable to establish their population. In the laboratory experiments, beside the growth of head capsule width between each larval instar corresponded to Dyar''s rule, the growth of the head capsule length and the length and width of prontum were examined to correspond to Dyar''s rule. The larval instar could be recognized by the morphological difference among each larval instar and the character that the growth of length and width of head capsule and prontum correspond to Dyar''s rule. In the growth experiment with 12L:12D photoperiod at 25℃, each of the developmental time from 1st to 6th instar larvae were 26.1 ± 0.6, 25.0 ± 1.1, 36.3 ± 1.2, 45.7 ± 2.6, 61.3 ± 2.5, and 52.4 ± 6.9 days, respectively. On the food preference of the 6th instar larvae, the larvae collected from rice field in Nan-Kang preferred the tadpole of Rana latouchii to freshwater snail of Sinotaia quadrata and fish of Rhinogobius brunneus. The larvae collected from two places showed no significant differences in feeding proportion among six freshwater snail species without shell. The shell width and length of the freshwater snails caught by the 6th instar larvae were from 1.6 to 2.4 mm and 2.8 to 10.9 mm, respectively. In another experiment, larvae showed no feeding preference among five species of freshwater snails. The factors that effected the larvae preyed on the freshwater snails depended on the body size of snails rather than the species.

壹、前言............................................................................................ 1
貳、前人研究.................................................................................... 3
參、材料與方法
一、野外試驗部份
（一）、試驗地點........................................................................... 11
（二）、試驗地點環境調查........................................................... 13
（三）、野放蟲源........................................................................... 14
（四）、族群估算........................................................................... 14
二、室內試驗部份
（一）、供試蟲源............................................................................ 16
（二）、黃緣螢幼蟲之飼養觀察.................................................... 16
（三）、幼蟲食物偏好性................................................................ 17
肆、結果與討論
一、野外試驗結果
（一）、試驗地點的環境調查結果................................................ 22
（二）、幼蟲存活數量.................................................................... 23
（三）、成蟲發生數量 ................................................................ 24
（四）、陸棲螢火蟲的發生............................................................ 26
二、室內試驗結果
（一）、黃緣螢卵及幼蟲形態........................................................ 28
（二）黃緣螢幼蟲發育時間............................................................ 31
（三）、黃緣螢幼蟲對食物種類的偏好性.................................... 33
伍、黃緣螢棲地經營管理問題及建議事項
一、野放經過及棲地經營管理過程................................................ 42
二、棲地經營管理問題及討論........................................................ 46
陸、結論............................................................................................ 50
柒、致謝............................................................................................ 52
捌、引用文獻.................................................................................... 53
表........................................................................................................ 62
圖........................................................................................................ 71
附錄一................................................................................................ 98
附錄二................................................................................................ 99

Annandale, N. 1900. Observation on the habitats and natural surroundings of Insecta made on the "SKeat Expedition" to the Malay Peninsula 1899-1900. VI. Insect luminosity: An Aquatic Lampyridae. Proc. Zool. Soc. London: 862-865. ( In Latin)
Annandale, N. 1906. Notes on the freshwater fauna of India, No. III. An Indian cockroach and beetle larva. J. Asiat. Soc. Bengal, 2, Calcutta, 1906: 105-108(cf. 106) (From Bertrand''s note, 1973)
Ballantyne, L. A., and E. Buck. 1979. Taxonomy and behavior of Luciola (Luciola) aphrogeneia, a new surf firefly from Papua New Guinea. Trans. Amer. Ent. Soc. 105-137.
Bernays, E. A., and R. F. Chapman. 1994. Host-Plant Selection by Phytophagous Insects. Chapman & Hall, New York. p. 219-226.
Bertrand, H. P. I. 1973. Larvae and pupae of water beetles collected from the island of Ceylon. Bull. Fish. Res. Stn., Sri Lanka (Ceyoln) 24: 95-112.
Beutel, R. G. 1995. Phylogenetic analysis of Elateriformia (Coleoptera: Polyphaga) based on larval characters. J. Zool. Syst. Evol. Research 33: 145-171.
Blair, K. G. 1927. An aquatic lampyrid larva from S. Celebes. Trans. Entomol. Soc., London, p. 43.(From Okada''s note, 1928)
Blum, M. S. and A. Sannasi. 1974. Reflex bleeding in the lampyrid Photinus pyralis: defensive function. J. insect Physiol. 20: 451-460.
Branham, M. A., and M. D. Greenfield. 1996. Flashing males win mate success. Nature 381: 745-746.
Buck, J., and J. F. Case. 1961. Control of flashing in fireflies. I. The lantern as a neuroeffector organ. Biol. Bull. 121: 234-256.
Buck, J., J. F. Case, and F. E. Hanson, jr. 1963. Control of flashing in fireflies. III. Peripheral excitation. Biol. Bull. 125: 251-269.
Buschman, L. L. 1984. Biology of the firefly Pyractomena lucifera (Coleoptera: Lampyridae). Fla. Entomol. 67(4): 529-542.
Bushman, L. L. 1988. Light organs of immature fireflies (Coleoptera: Lampyridae) as eye-spot / false- head desplays. Coleopt. Bull. 42(1): 94-97.
Carlson, A. D., and J. Copeland. 1972. Photic inhibition of brain stimulated firefly flashing. Am. Zool. 12: 479-487.
Case, J. F. 1984. Vision in mating behaviour of fireflies. p. 195-222. in T. Lewis, ed. Insect Communication. Academic Press, New York.
Case, J. F., and J. Buck. 1963. Control of flashing in fireflies. II. Role of central nervous system. Biol. Bull. 125: 234-250.
Chang, J. C. 1994. Studies on artificial rearing of aquatic firefly, Luciola ovalis Hope, in Taiwan. Master thesis, Dept. Entomology of National Chung-Hsing University, Taichung. 48 pp. ( In Chinese)
Chang, J. C., and P. S. Yang. 1998a. The species and ecology of the fireflies in the national parks. The Ecology and Conservation of the Fireflies Symposium for National Parks p. 65-72. ( In Chinese)
Chang, J. C., and P. S. Yang. 1998b. The habitat of the fireflies. The Ecology and Conservation of the Fireflies Symposium p. 65-67. (In Chinese)
Chang, J. C., and P. S. Yang. 1998c. The breeding and observation of the fireflies. The Ecology and Conservation of the Fireflies Symposium p. 68-73. (In Chinese)
Chang, W. S. 1998. Building a artificial breeding environment for the fireflies. The Ecology and Conservation of the Fireflies Symposium p. 163-172. (In Chinese)
Cody, R. P., and J. K. Smith. 1997. Applied Statistics and the SAS Programming Language (Fourth Edition). Prentice-Hall, Upper Saddle River, New Jersey. 445 pp.
Crawford S. C. 1934. The habits and characteristic of nocturnal animals. Q. Rev. Biol. 9: 201-214. (From Blum and Sannasi''s note, 1973)
Crowson, R. A. 1972. A review of the classification of Cantharoidea (Coleoptera), with the definition of two new families, Cneoglossidae and Omethidae. Revista de la Universidad de Madrid 21: 35-77.
De Boer, G., and F. E. Hanson. 1984. Food plant selection and induction of feeding preference among host and non-host plants in larvae of the tobacco hornworm Manduca sexta. Entomol. exp. appl. 35: 177-193.
Eguchi, E., A. Nemoto, V. B. Meyer-Rochow, and M. Ohba. 1984. A comparative study of spectral sensitivity curves in three diuranal and eight nocturnal species of Japanese fireflies. J. Insect Physiol. 30: 607-612.
Eisner, T., D. F. Wiemer, L. W. Haynes, and J. Meinwald. 1978. Lucibufagins: Defensive steroids from the fireflies Photinus ignitus and P. marginellus (Coleoptera: Lampyridae). Proc. Natl. Acd. Sci. 75: 905-908.
Eisner, T., M. A. Goetz, D. E. Hill, S. R. Smedley, and J. Meiwald. 1997. Firefly "femmes fatles" acquire defensive steroids (lucibufagins) from their firefly prey. Proc. Natl. Acad. Sci. 94: 9723-9728.
Frost, S. W. 1959. Chapter IX. Color. p.176-197. in: Insect Life and Insect Natural History. Dover Publications, Inc. New York. 526pp.
Furuta, T. 1991. A report of activeties for restoration of firefly environment in the educational program in a middle school. p. 95-101. in: Reconquista SP. No.1. Bulletin of the Natural Environmental Restoration. Saiteku Press, Tokyo.136 pp.（In Japanese）
Gall, L. F. 1985. Measuring the size of lepidopteran populations. J. Res. lepid. 24: 97-116.
Haneda, Y. 1977. The experiment of artificial in fireflies. Bull. Yokosuka City Mus. 23: 23-26.( In Japanese)
Haneda, Y. 1985. Luminous Organisms. Kosei Press, Tokyo. 318 pp.( In Japanese)
Hanson, W. R. 1967. Estimating the density of an animal population. J. Res. Lepid. 6: 203-247.
Harvey, E. N., and R. T. Hall. 1929. Will the adult firefly luminesce it its larval organs are entirely removed? Science 69: 253-254.
Ho, C. Z., and B. H. Chiang. 1997. Two firefly species with aquatic larvae in Taiwan. Conservation Quarterly 17: 42-46. ( In Chinese)
Ho, C. Z., C. S. Chu, and C. C. Chu. 1998. A discovery of the aquatic larvae of Laciola[sic!] substriata. Nature Conservation Quarterly22: 47-51. ( In Chinese)
Hsia, W. T., and S. S. Kao. 1985. The meanings of head width measurements for larvae in Entomology. Scientific Agriculture 33: 285-289.( In Chinese)
Imai, H. 1955. On the chromosomes in some Lampyridae. Japan. J. Genet. 30: 169.(from Virkki''s note, 1963)
Jaenike, J. 1988. Effects of early adult experience on host selection in insects: some experimental and theoretical results. J. Insect Behav. 1: 3-15.
Jeng, M., J. Lai, and P. Yang. 1999. A synopsis of the firefly fauna at six national parks in Taiwan (Coleoptera: Lampyridae). Chinese J. Entomol. 19: 65-91.( In Chinese)
Jeng, M., P. Yang, and M. Sato. 1998a. The genus Cyphonoderus (Coleoptera, Lampyridae) form Taiwan and, with notes on the subfamily Cyphonoderinae. Elytra 26: 379-398.
Jeng, M., P. Yang, M. Sato, J. Lai, and J. Chang. 1998b. The genus Curtos (Coleoptera, Lampyridae, Luciolinae) of Taiwan and Japan. Japan J. Syst. Entomol. 4: 331-347.
Kanda, 1935. Firefly. The Bulletin of Luminous Organisms, Tokyo. 496 pp.( In Japanese)
Klingenberg, C. P. 1992. Dyar''s rule and multivariate allometric growth in nine species of waterstriders (Heteroptera: Gerridae). J. Zool. 227: 453-464.
Kobayashi, H. 1991. A report of restoration of natural environment for firefly from Shiojiri City. p. 65-73. in: Reconquista SP. No.1. Bulletin of the Natural Environmental Restoration. Saiteku Press, Tokyo.136 pp. ( In Japanese)
Kondo, A., and F. Tanaka. 1989. An experimental study of predation by the larvae of the firefly, Luciola lateralis Motshulsky (Coleoptera: Lampyridae) on the apple snail, Pomacea canaliculata Larmarck (Mesogastropoda: Pillidae). Jpn. J. Appl. Entomol. Zool. 33: 211-216. ( In Japanese)
Krebs, C. J. 1994. Chapter 10: Population parameters. p. 151-167. in: Ecology: The Experimental Analysis of Distribution and Abundance (4th). HarperCollins College Publishers, New York.
Kuan, C. 1984. The Luminescence of insects. Bulletin of the Society of Entomology. 17: 51-58.（In Chinese）
Kuon, K. S. 1991. Entomology. Vol. 1. (reedition). Cow College of National Chung Hsing University, p. 237-238.( In Chinese)
Lai, J. 1998. Revision of Lampyrinae from Taiwan (Coleoptera: Lampyridae). Master''s thesis, Graduate Institute of Plant Pathology & Entomology , National Taiwan University. 120 pp.
Lai, J., M. Sato, and P. Yang. 1998. Checklist of Lampyridae of Taiwan - Coleoptera: Polyphaga: Lampyridae. Chinese J. Entomol. 18: 207-215. ( In Chinese)
Lall, A. B. 1993. Action spectra for the initiation of bioluminescent flashing activity in males of twilight-active firefly Photinus scintillans (Coleoptera: Lampyridae). J. Insect Physiol. 39: 123-127.
Lall, A. B., G. K. Strother, T. W. Cronin, and H. H. Seliger. 1988. Modification of spectral sensitivities by screening pigments in the compound eyes of twilight-active fireflies (Coleoptera: Lampyridae). J. Comp. Physilo. A 162: 23-33.
Lloyd, J. E. 1965. Aggressive mimicry in Photuris: firefly femmes fatales. Science 149: 653-654.
Lloyd, J. E. 1968. Illumination, another function of firefly flashes? Entomol. News 79: 265-268.
Lloyd, J. E. 1969. Flashes of Photuris fireflies: their value and use in recognizing species. Fla. Entomol. 29-35.
Lloyd, J. E. 1971. Bioluminescent communication in insect. Ann. Rev. Entomol. 16: 97-122.
Lloyd, J. E. 1973a. A firefly inhabitant of coastal reefs in New Guinea (Coleoptera: Lampyridae). Biotropica 5: 168-174.
Lloyd, J. E. 1973b. Firefly parasites and predators. Coleopt. Bull. 27: 91-106.
Lloyd, J. E. 1983. Bioluminescence and communication in insects. Ann. Rev. Entomol. 28: 131-160.
Lloyd, J. E., and S. R. Wing. 1989. Flash behavior and ecology of Thai Luciola fireflies (Coleoptera: Lampyridae). Fla. Entomol. 72(1): 80-85.
McDermott, F. A. 1910. A note on the light-emission of some American Lampyridae. Can. Entomol. 42: 357-363.(from Blum and Sannasi''s note, 1973)
McDermott, F. A. 1953. Glow-worms in a marine littoral habitat in Jamaica (Coloeptera; Lampyridae). Entomol. News 64: 89-90.
McDermott, F. A. 1964. The taxonomy of the Lampyridae (Coleoptera). Trans. Am. Entomol. Soc. 90: 1-72.
McDermott, F. A. 1966. Coleopterorum Catalogus, Supplementa 9, Lampyridae. N. V. Drukkerij Hooiberg, Netherlands. 149 pp.
Meinwald, J., D. F. Wiemer, and T. Eisner. 1979. Lucibufagins. 2. Esters of 12-oxo-2β,5β,11α-trihydroxybufalin, the major defensive steroids of the firefly Photinus pyralis (Coleoptera: Lampyridae). J. Am. Chem. Soc. 101: 3055-3060.
Miishi, T.。1996. Luciola lateralis. Hoozuki Press. 241 pp. ( In Japanese)
Miyashita, M. 1991. Revival of fireflies in rice field. p. 83-93. in: Reconquista SP. No.1. Bulletin of the Natural Environmental Restoration. Saiteku Press, Tokyo.136 pp. ( In Japanese)
Mori, S. 1991. Reconstruction of the natural waterside for the revival of firefly. p. 35-48. in: Reconquista SP. No.1. Bulletin of the Natural Environmental Restoration. Saiteku Press, Tokyo.136 pp. (In Japanese)
Murakami, M. 1991. A history of firefly preservation activities in Japan. p. 23-32. in: Reconquista SP. No.1. Bulletin of the Natural Environmental Restoration. Saiteku Press, Tokyo.136 pp. ( In Japanese)
Nagai, S., H. Yamamoto, K. Ishii, M. Otuka, and M. Kobayashi. 1979. Rearing and population growth of fresh water snail, Semisulcospira libertina in the laboratory. VENUS (Japan. J. Malac.) 38: 25-34.
Nakane, T. 1991. Lampyrid insects of the world. p.3-11. in: Reconquista SP. No.1. Bulletin of the Natural Environmental Restoration. Saiteku Press, Tokyo.136 pp. ( In Japanese)
Nobi Firefly Research Club. 1990. Ecological research of the firefly and analysis of its habitats, Nobi area, Yokosuka City, Kanagawa Prefecture. Sci. Rept. Yokosuka City Mus. 38: 47-60.( In Japanese)
Ogata, T. 1991. The movements of reconstruction of natural environment for firefly in Kumanoto Prefecture. p. 51-63. in: Reconquista SP. No.1. Bulletin of the Natural Environmental Restoration. Saiteku Press, Tokyo.136 pp. ( In Japanese)
Ohba, N. 1973. Bleeding process of Luciola cruciata and L. lateralis in small water tank. Bull. Yokosuka City Mus. 18: 1-5.( In Japanese)
Ohba, N. 1974. Bleeding process of Luciola cruciata and L. lateralis in small water tank (2). Bull. Yokosuka City Mus. 19:1-7 ( In Japanese)
Ohba, N. 1977a. The influence of the chemicals to the first instar of Luciola cruciata Motschulsky. Bull. Yokosuka City Mus. 23: 14-15. (In Japanese)
Ohba, N. 1977b. The artificial food of Luciola cruciata and L. lateralis. Bull. Yokosuka City Mus. 23: 16-20. ( In Japanese)
Ohba, N. 1981. Luminous insect and museums in Taiwan. Bull. Yokosuka City Mus. 27: 43-46. ( In Japanese)
Ohba, N. 1983. Studies on the communication system of Japanese fireflies. Science Report of the Yokosuka City Museum No.30. 62 pp.
Ohba, N. 1986. The ecology and habitat of Luciola cruciata at Nobi District in Yokosuka City. Bull. Yokosuka City Mus. 33: 1-2. ( In Japanese)
Ohba, N. 1988. Aquatic fireflies. The Nature and Insects 23: 8-13. ( In Japanese)
Ohba, N. 1989. The differences between the fireflies, Luciola cruciata, in western Japan and eastern Japan. The Nature and Insects 24: 2-6. ( In Japanese)
Ohba, N. 1993. Breeding and Observation of the Fireflies. Hot Press, Tokyo. 167 pp.( In Japanese)
Ohba, N., T. Tsumuraya, K. Honda, S. Murata, and Y. Omori. 1993. Ecological study of the firefly, Luciola lateralis, of the Kushiro Shitsugen and Akkeshi, Hokkaido. Sci. Rept. Yokosuka City Mus. 41: 15-26. ( In Japanese)
Ohba, N., S. Azuma, K. Nishiyama, Y. Goto, H. Suzuki, Y. Sato, and I. Kawashima. 1994. Morphology, life history and behaviour of the firefly, Luciola owadai (Coleoptera: Lampyridae). Sci. Rept. Yokosuka City Mus. 41: 15-26. ( In Japanese)
Okada, Y. K. 1928. Two Japanese glow-worms. Trans. Entomol. Soc. London 76: 101-108.
Ozawa, T. 1991. Special plan and techniques of embankment for revival of firefly. p. 75-82. in: Reconquista SP. No.1. Bulletin of the Natural Environmental Restoration. Saiteku Press, Tokyo.136 pp. ( In Japanese)
Sato, M., and M. Kimura. 1994. Discovery of a new firefly of the Genus Luciola (Coleoptera, Lampyridae) from Kume jima of the Ryukyu Island. Elytra 22: 159-162.
Schwalb, H. H. 1961. Beitrage zur biologie der einheimischen Lampyriden Lampyris noctiluca Geoffr. und Phausis splendidula Lec. und experimentelle analyse ihres Beutefang- und sexualverhaltens. Zool. Jb. Syst. Bd. 4: 399-550.
Sheppard, P. N., and J. A. Bishop. 1973. The study of populations of Lepidoptera by capture-recapture methods. J. Res. Lepid. 12: 135-144.
Shibue, K., N. Ohba, and E. Fujii. 1995. Analysis of habitat factor which influence on population of Luciola cruciata at Nobi District in the Miura Peninsula. J. JILA 58:121-124. ( In Japanese)
Sivinski, J. 1981. The nature and possible functions of luminescence in Coleoptera larvae. Coleopt. Bull. 35: 167-179.
Southwood, T. R. E. 1978. Chapter 3: Absolute population estimates using marking techniques. p. 70-129. in: Ecological method, with particular reference to the study of insect populations(2nd). Chapman & Hall Press, London.
Sydow, S. L., and J. E. Lloyd. 1975. Distasteful fireflies sometimes emetic, but not lethal. Fla. Entomol. 58: 312.
Taylor, R. L. 1931. On "Dyar''s Rule" and its application to sawfly larvae. Ann. Entomol. Soc. Am. 24: 451-466.
Underwood, J. T., W. D. Tallamy, and J. D. Pesek. 1997. Bioluminescence in firefly larvae: a test of the aposematic display hypothesis (Coleoptera: Lampyridae). J. Insect Behav. 10(3): 368-370.
Virkki, N. 1963. On the cytology of some neotropical cantharoids (Coleoptera). Ann. Acad. Sci. Fenni. IV Biologica 65: 1-16.
Water System Research Club. 1989. Recovery of the urban water system, Iwato River, Yokosuka, Kanagawa Prefecture. Sci. Rept. Yokosuka City Mus.37: 9-28. ( In Japanese)
Wey, Y. S. 1995. Ecological and habit studies of adult blue tigers (Lepidoptera: Nymphalidae, Danainae) in Tatun Mountains. Dissertation thesis, Graduate Institute of Plant Pathology and Entomology of National Taiwan University. 171 pp. ( In Chinese)
Wey, Y. S., and P. S. Yang. 1990. Evaluation of lepidopteran populations: mark-release-recapture models. Taipei Zoo Bull. 2: 119-131. ( In Chinese)
Winston, L. M. 1987. The Biology of the Honey Bee. Harvard University Press, London. p. 176.
Wu, C. F. 1937. Super family Cantharoidea: Family Lampyridae. Catologus Insectorum Sinensium 3: 377-385.
Wu, H. 1998. Taxonomy of Calamoceratidae and Lepidostomatidae (Trichoptera: Integripalpia) in Taiwan. Master''s thesis, Graduate Institute of Plant Pathology & Entomology , National Taiwan University. 127 pp. ( In Chinese)
Yajima, M., and N. Ohba. 1996. The fireflies handbook. 72 pp.（In Japanese）
Yang, P. S. 1998. The study of the fireflies in Taiwan in the past and future. The Ecology and Conservation of the Fireflies Symposium p. 1-9. ( In Chinese)
Yuma, M. 1984. Egg size and viability of the firefly, Luciola cruciata (Coleoptera, Lampyridae). Kontyu 52: 615-629.
Yuma, M., and M. Hori. 1990. Seasonal and age-related changes in the behaviour of the Genji firefly, Luciola cruciata (Coleoptera, Lampyridae). Jpn. J. Entomol. 58: 863-870.