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研究生:林靖惠
研究生(外文):Lin Jinghuei
論文名稱:棲蘭山台灣樹繁殖枝條葉部性狀之研究
論文名稱(外文):Leaves Attributes of Reproductive shoots of Taiwan Sassafras(Sassafras randaiense Rhed.) at Chilan Shan,Northeastern Taiwan
指導教授:關秉宗林世宗林世宗引用關係
指導教授(外文):Dr.Biing T.GuanDr.Shu T.Lin
學位類別:碩士
校院名稱:國立臺灣大學
系所名稱:森林學研究所
學門:農業科學學門
學類:林業學類
論文種類:學術論文
論文出版年:2000
畢業學年度:88
語文別:中文
論文頁數:67
中文關鍵詞:台灣樹冠層空間葉綠素指標值混合模式Mixed ModelSPAD-502
外文關鍵詞:Sassafras randaienseHay.Rehdercanopy regionchlorophyll concentrationMixed ModelSPAD-502
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台灣樹(Sassafras randaiense(Hay.)Rehder)為台灣落葉性特有闊葉樹種,經評估為易受害(vulnerable)之稀有植物,其亦為寬尾鳳蝶的主要食草,因此在生物及生態上的地位特殊,極具保育價值。本研究針對台灣樹冠層空間葉部形態和葉綠素指標值隨時間推移的變化,探討台灣樹的調適能力。
本研究以棲蘭山神木園區內台灣樹為調查對象,於民國八十七年七月至十月間,平均每十四天量測樣木不同冠層部位之葉長、寬,並以SPAD-502測計同一片葉的葉綠素指標值。利用重複觀測的層級(nested)試驗設計,第一層級為單木,次層級則為冠層層次與部位,重複觀測六次。所得結果以SAS混合模式(Mixed Model)程序(Proc Mixed),進行變異分析,求出葉綠素指標值與葉部形態於不同林木位置之樣木間及其冠層空間位置上之變化,以判釋各參數在與時間及空間上的相關性。結果顯示台灣樹葉部性狀在樣木間的效應顯著;台灣樹冠層空間內葉綠素指標值及葉面積的變化在不同樹冠部位的差異顯著,而層次上的差異則不明顯。台灣樹冠層空間時間效應分析上,葉面積之變異大,主要受葉長影響,但自我相關係數隨時間延長而相對降低。葉綠素指標值在十月上旬時變異增大,應與落葉現象有關。
Taiwan sassafras (Sassafras randaiense Rehder.), a native deciduous broad-leaved specie, is the sole food source for caterpillars of the endangered broad-tailed swallowtail butterfly (Agehana maraho). The tree species is also being regarded as a vulnerable species by some researchers. As a part of an ongoing project to understand the population biology and ecology of this species, this study focused on the dynamics of foliar attributes throughout a growing season. The attributes examined included leaf chlorophyll concentration (measured by Minolta SPAD-502 chlorophyll meter), leaf length, leaf width, and leaf size.
Representing different shading conditions, five trees located at Chilan Shan, northeastern Taiwan, were selected for this study. The canopy of each tree was divided into nine regions; each region was a combination of a relative vertical position (top, middle, or bottom) and a relative horizontal position (outer, central, or inner). For each canopy region, 20 leaves growing on reproductive shoots were subjectively selected and repeatedly measured for six times (from July to October of 1998). The experiment design was thus a repeated-measure nested design with individual tree serving as the primary unit and each canopy region as the secondary unit. A mixed model ANOVA was used to analyze the data.
The results showed that chlorophyll concentration, leaf width and leaf size differed significantly among individual trees; trees with an isolated canopy had higher chlorophyll concentration, wider and larger leaves during the observation period. The four attributes did not differ significantly alone the vertical direction. However, the attributes did differ significantly alone the horizontal direction. Leaves in the outer and central parts of the canopy had higher chlorophyll concentration and were longer, wider and bigger. In general, the variability of the four attributes increased throughout the growing season, with the last measurement having the greatest variance. The beginning of translocation of chlorophylls by some leaves at the last measurement led to the largest variance in chlorophyll concentration. The four attributes also showed different variation patterns through time. Consecutive measurements of leaf chlorophyll concentration and leaf width did not have a significant correlation. Consecutive measurements of individual leaf length and leaf size, however, had large and significant autocorrelation coefficients, and the autocorrelations decreased with increasing measurement lags.
謝誌
中文摘要
英文摘要
壹、前言
貳、前人研究
(一)樹之研究暨物候觀察概述
(二)冠層結構與葉片空間分佈對林木之影響
(三)葉片特徵與環境之關係
(四)葉的生長期和光環境之關係
參、調查材料與方法
(一)研究地概況
(二)調查方法
(三)理論基礎暨分析方法
肆、結果
(一)台灣樹葉綠素含量變化
(1)樣木效應
(2)冠層部位效應
(3)時間效應
(二)台灣樹葉形態上的變化
(1)樣木效應
(2)冠層部位效應
(3)時間效應
伍、討論
陸、結論
柒、參考文獻
附錄一、混合模式之SAS程式
附錄二、樣木及部位間葉綠素指標值之分析結果
附錄三、樣木部位間葉形態比較結果
附錄四、研究調查照片
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