(18.210.12.229) 您好!臺灣時間:2021/03/05 12:59
字體大小: 字級放大   字級縮小   預設字形  
回查詢結果

詳目顯示:::

我願授權國圖
: 
twitterline
研究生:邱美倫
研究生(外文):Chiou Mei-Luen
論文名稱:台灣海域產鼠尾鱈科魚類之系統分類研究
論文名稱(外文):Systematic Studies on the Macrourid Fishes (Family Macrouridae,Gadiformes)from the Waters Around Taiwan.
指導教授:沈世傑沈世傑引用關係邵廣昭邵廣昭引用關係
指導教授(外文):Shen Shih-ChiehShao Kwang-Tsao
學位類別:碩士
校院名稱:國立臺灣大學
系所名稱:動物學研究所
學門:生命科學學門
學類:生物學類
論文種類:學術論文
論文出版年:2001
畢業學年度:90
語文別:中文
論文頁數:85
中文關鍵詞:系統分類學鼠尾鱈科深海魚
外文關鍵詞:SystematicMacrouridae
相關次數:
  • 被引用被引用:2
  • 點閱點閱:478
  • 評分評分:系統版面圖檔系統版面圖檔系統版面圖檔系統版面圖檔系統版面圖檔
  • 下載下載:0
  • 收藏至我的研究室書目清單書目收藏:0
摘要
鼠尾鱈(Family Macrouridae)為深海底棲性魚類,多數生活在溫帶及熱帶海域水深200~6000公尺之間,全世界共4亞科32屬約300種(Nelson,1994),1995至1999年間則又陸續發表26種。目前台灣海域水深200~800公尺之間共採獲有2亞科11屬40種,其中包括腔吻鱈屬(Caelorinchus) 的3種新種窄吻腔吻鱈(Caelorinchus leptorhinus)、鞍斑腔吻鱈(C.ephippins)與長絲腔吻鱈(C.logispinosus),及台灣地區1個新紀錄亞科、6個新記錄屬、19個新記錄種。分別為新紀錄底尾鱈亞科(Bathygadinae)之鞭鼠鱈(Gadomus colletti)、 孔頭底尾鱈(Bathygadus antrodes)、日本底尾鱈(B.nipponicus)、鬚底尾鱈 (B.garretti);以及鼠尾鱈亞科(Macrourinae)之細身膜鱈(Hymenogadus gracilis)、刺吻膜首鱈(Hymenocephalus lethonenus)、獅鼻奈氏鱈(Nezumia condylura)、偽鯨尾鱈(Pseudoctenurous septifer)、日本舟尾鱈 (Kumba japonica)、長鬚凹腹鱈(Ventrifossa longibarbata)、大鰭凹腹鱈(V.macroptera)、扇鰭凹腹鱈(V.rhipidodorsalis)、西海凹腹鱈(V.saikaiensis)、魔燈凹腹鱈(V.lucifer)、歧異凹腹鱈(V.divergens)、箭齒凹腹鱈(V.atherodon)、東海腔吻鱈(Caelorinchus productus)、長管腔吻鱈(C.longissimus)及擬星腔吻鱈(C.asteroides)等。
由於深海魚類採集技術較為困難,且採獲之標本多半殘破不堪、難以正確鑑別屬種、所以過去台灣的鼠尾鱈科魚類研究較為欠缺,另外舊有的記錄中大部份種類亦無標本可供查考,因此本研究為過去文獻中缺乏標本的種類,重新提供登錄的標本證據,亦修訂了早期報告中誤鑑、同種異名及位階安置不當等問題。文中除整理出鼠尾鱈科魚類各亞科、屬及種撿索表、同種異名錄、鑑別特徵、體型、分佈,相似種間之比較和備註等資料外,並附以台灣產40種鼠尾鱈的彩色標本照以供相關研究人員分類鑑定之參考。
在類緣關係方面,利用16個測量形質、8個計數形質及67個其它一般外部形質等91個特徵,以表型及支序分析方法推斷其類緣關係。其結果在鼠尾鱈亞目中鼠尾鱈科,包含各亞科及若干屬與亞屬間的關係與過去所作之研究結果並不一致。表型聚類分析與支序分析結果顯示底尾鱈亞科、鼠尾鱈亞科、Caelorinchus屬與Hymenogadus-Hymenocephalus group各為4組相當獨立的單系群。支序分析結果顯示底尾鱈亞科為最原始的一群而Caelorinchus屬為最特化的一群。Ventrifossa屬在表型分析中聚為一單系群,但在支序分析中各種間關係混淆,Hymenogalus-Hymencephalus group 在表型分析中之差異達屬級水準,但在支序分析中未達屬級水準; Caelorinchus屬4亞屬間關係在2種分析結果均混淆,因此本研究不贊同將Caelorinchus屬區分成4亞屬,Hymenogadus屬與Hymenocephalus屬在表型分析與支序分析有不同的結果,因此兩者間的差異是否達到屬級水準,則有待更深入的探究。Kumba屬、Pseudoctenurous屬、Malacocephalus屬在表型分析中類緣關係較接近Ventrifossa屬;在支序分析中Pseudoctenurous屬與Malacocephalus屬較接近,Kumba屬關係混淆。Nezumia屬與Coryphaenoides屬在表型分析中為關係接近的2群,但在支序分析中關係混淆。
在地理分佈方面,台灣周邊海域以東北海域大溪之魚類相最為豐富,目前的資料亦顯示台灣東北部及西南部的種類組成有明顯差異;在豐度方面,鼠尾鱈科魚類中則以Caelorinchus kishnouyei、C.multispinulosus、C.formosanus和Ventrifossa rhipidodorsalis的分佈最廣,數量也最多。在印度太平洋區所有的233種鼠尾鱈科魚類中則以台灣、中國、日本三者的鼠尾鱈魚種組成最為接近,其次為菲律賓及東印度群島,66%種類目前只發現在單一區域,6%的種類分佈超過3個區域為本科魚類中少見之廣泛種,此顯示鼠尾鱈的稀有種比例高,其原因可能是採樣仍不足,或多數鼠尾鱈之地理分佈範圍並不廣泛之故。
Abstract
The grenadiers (Family: Macrouridae) is one of the major group of benthopelagic fishes. Grenadiers occupy all oceans, except the high Arctic, and almost all basins with deep oceanic connections; none normally occur in depths shallower than found at abyssal depths. One species to more than 6000m, most species are found between 200m and 2000m (Iwamoto, 1990). At present, Grenadiers are approximately divided into 4 subfamilies, 38 genera and 327 species in the world. According to "Fishes of Taiwan" (Shen et al, 1993), there were 1 subfamily, 5 genera and 19 species occurred in Taiwan. The present study attempted to review this group of fish by re-examining previous deposited specimens and re-collecting new specimens from deep sea bottom trawler fisheries from 300-800 meters in north-eastern and south-western waters near Tashi, Nanfanao and Tungkang . Our preliminary study results obtained a total number of 2 subfamilies,11 genera and 40 species in which 19 species are new records and 3 are new species .They include three new species of Caelorinchus leptorhinus ,C.ephippins and C.logispinosus, and new records of 19 species: Gadomus coletti, Bathygadus antrodes, B.nipponicus, B.garretti, Hymenogadus gracilis, Hymenocephalus lethonenus, Nezumia condylura, Pseudoctenurous septifer, Kumba japonica, Ventrifossa longibarbata , V.macroptera, V.rhipidodorsalis V.saikaiensis, V.Lucifer, V.divergens, V.atherodon, C.productus, C.longissimus and C.asteroids.
Due to the difficulty and the divergence of character’s identification, the classification of this group of fish have many synonymous, misidentification and different classification system problems. The diagnostic key and the specimen color photos of each species are provided.
Additionally, the phylogenetic relationship of these 11 genera and 40 species were constructed using both phenetic and cladistic methods. Using 91 morphological characters to construct the phylogenetic relationships by phenetic (NTSYS) and cladistic (Henning 86).In conclusion:the subfamily Bathygadinae, Macrouridae, genus Caelorinchus and Hymenogadus-Hymenocephalus group are surely four monophyletic groups.The cladistic analysis shows Bathygadinae is more primitive than other taxa and Caelorinchus is the most specialize group for Macrouridae. Macrourinae and Bathygadinae are sister group. The relationship for Ventrifossa is a monophyletic group ; Hymenogadus and Hymenocephalus reach the genus level; Nezumia and Coryphaenoides are sister group; Kumba, Pseudoctenurous, Malacocephalus and Ventrifossa are very close in phenetic analysis. The phylogeny of Ventrifossa is not a monophyletic group ; Hymenogadus and Hymenocephalus not reach the genus level; Kumba ,Pseudoctenurous , Malacocephalus, Nezumia and Coryphaenoides are still unresolved in cladistic analysis. The difference for Oxymacrurus, Oxygadus, Paramacrurus and Quincunica of Caelorinchus are not reach the subgenus level in phenetic and cladistic analysis. So in the present study that the rank of subgenus for Caelorinchus differ from the generally-acknowledged system.
The thesis also analysis the distribution and abundance data of macrourids from the waters around Taiwan. Tashi has the highest fish fauna compare with the Tungkang and Nanfanao. Caelorinchus formorsanus , C.kishnouyei, C.multispinulosus and Ventrifossa rhipidodorsalis are the dominant species regarding to the total catch. As to the zoogeographical distribution, the species composition of grenadiers between north and south Taiwan are quite different. This result is quite consistent with the pattern obtained from coral reef and estuarine fishes. It may imply that the different fish fauna between northern and southern Taiwan can extend to the fishes which inhabit in deep waters. The thesis review the macrourids fauna of 9 areas of Indo-Pacific region. Taiwanese macrourids fauna are most similar to that of Japan and China. The global geographical distribution of macrourid species should be affected by the factors of degree of latitude, water temperature, ocean currents, land barrier, geographical distance, depth and substrata.
目次
謝辭 ---------------------------------------------------------i
中文摘要 -----------------------------------------------------ii
英文摘要 -----------------------------------------------------iv
壹、 前言------------------------------------------------------1
貳、 材料與方法------------------------------------------------5
一、 材料--------------------------------------------------5
(一) 標本的收集---------------------------------------5
(二) 標本的處理---------------------------------------5
二、 方法--------------------------------------------------5
(一) 外部形態的測定-----------------------------------5
1.測量形質方面-----------------------------------5
2.計數形質方面-----------------------------------7
3.其他一般外部形質方面---------------------------7
(二) 類緣關係之推斷-----------------------------------8
1. 資料矩陣的構成---------------------------------8
(1) 形質種類------------------------------------8
(2) 資料轉碼------------------------------------8
2. 表型聚類分析-----------------------------------8
3. 支序分析---------------------------------------9
(三) 地理分佈之探討-----------------------------------9
參、 結果-----------------------------------------------------10
一、 分類描述---------------------------------------------10
鼠尾鱈科Family Macrouridae---------------------------10
鼠尾鱈科之亞科與屬之檢索表----------------------------11
底尾鱈亞科Subfamily Bathygadinae---------------------12
底尾鱈屬Bathygadus-----------------------------------12
底尾鱈屬之種檢索表-----------------------------------13
孔頭底尾鱈Bathygadus antrodes----------------------13
日本底尾鱈Bathygadus nipponicus--------------------14
鬚底尾鱈Bathygadus garretti------------------------15
鼠鱈屬Gadomus----------------------------------------16
鞭鼠鱈Gadomus colletti-----------------------------16
鼠尾鱈亞科Subfamily Macrourinae----------------------------------17
膜鱈屬Hymenogadus------------------------------------18
細長膜鱈Hymenogadus gracilis-----------------------18
膜首鱈屬Hymenocephalus-------------------------------19
膜首鱈屬之種檢索表-----------------------------------20
長頭膜首鱈Hymenocephalus longiceps----------------20
刺吻膜首鱈Hymenocephalus lethonemus---------------21
紋喉膜首鱈Hymenocephalus striatissimus------------22
偽鯨尾鱈屬Pseudoctenurous----------------------------23
偽鯨尾鱈Pseudoctenurous septifer-------------------23
奈氏鱈屬Nezumia--------------------------------------24
獅鼻奈氏鱈Nezumia condylura------------------------25
舟尾鱈屬Kumba----------------------------------------26
日本舟尾鱈Kumba japonica---------------------------26
凹腹鱈屬Ventrifossa----------------------------------27
凹腹鱈屬之種檢索表------------------------------------28
箭齒凹腹鱈Ventrifossa atherodon---------------------29
黑背鰭凹腹鱈Ventrifossa nigrodorsalis---------------29
加氏凹腹鱈Ventrifossa garmani-----------------------30
歧異凹腹鱈Ventrifossa divergens---------------------31
大鰭凹腹鱈Ventrifossa macroptera--------------------32
黑緣凹腹鱈Ventrifossa nigromarginta-----------------33
長鬚凹腹鱈Ventrifossa longibarbata------------------33
扇鰭凹腹鱈Ventrifossa rhipidodorsalis---------------34
西海凹腹鱈Ventrifossa saikaiensis-------------------35
魔燈凹腹鱈Ventrifossa lucifer-----------------------36
軟首鱈屬Malacocephalus--------------------------------36
滑軟首鱈Malacocephalus laevis-----------------------37
突吻鱈屬Coryphaenoides--------------------------------38
暗邊突吻鱈Coryphaenoides marginatus-----------------39
腔吻鱈屬Caelorinchus----------------------------------40
腔吻鱈屬之種檢索表-------------------------------------41
日本腔吻鱈Caelorinchus (Oxymacrurus) japonicus-------42
平棘腔吻鱈Caelorinchus (Oxygadus) parallelus---------43
胸斑腔吻鱈Caelorinchus (Paramacrurus) kishnouyei-----44
鴨嘴腔吻鱈Caelorinchus (Oxymacrurus) anatirostris----45
東海腔吻鱈Caelorinchus (Oxymacrurus) productus-------46
帶斑腔吻鱈Caelorinchus (Paramacrurus) cinguletus-----46
斯氏腔吻鱈Caelorinchus (Oxymacrurus) smithi----------47
吉氏腔吻鱈Caelorinchus (Oxygadus) gilberti-----------48
哈氏腔吻鱈Caelorinchus (Quincuncia) hubbsi-----------49
多棘腔吻鱈Caelorinchus (Quincuncia) multispinulosus--50
蒲原氏腔吻鱈Caelorinchus (Quincuncia) kamoharai------50
長管腔吻鱈Caelorinchus (Quincuncia) longissimus------51
擬星腔吻鱈Caelorinchus (Oxymacrurus) asteroides------52
臺灣腔吻鱈Caelorinchus (Quincuncia) formosanus-------52
窄吻腔尾鱈Caelorinchus (Oxymacrurus) leptorhinus-----53
鞍斑腔吻鱈Caelorinchus (Oxygadus) ephippins----------56
長絲腔吻鱈Caelorinchus (Quincuncia) logispinosus-----60
  二、類緣關係之推斷------------------------------------------63
(一) 表型分類法------------------------------------------63
1. 傳統測量形質方面----------------------------------63
2. 計數形質方面--------------------------------------63
3. 其他一般形質方面----------------------------------64
(二) 支序分類法------------------------------------------66
三、 地理分佈------------------------------------------------68
肆、 討論--------------------------------------------------------72
一、 分類描述------------------------------------------------72
二、 類緣關係------------------------------------------------74
(一)表型聚類分析------------------------------------------------------------76
1.測量形質方面--------------------------------------76
2.計數形質方面--------------------------------------76
3.其他一般形質方面----------------------------------77
(二)支序分析------------------------------------------------------------------78
三、 地理分佈------------------------------------------------80
伍、引用文獻----------------------------------------------------82
陸、附表
柒、附圖
捌、彩色圖版
引用文獻
沈世傑。1984a。台灣魚類檢索。南天書局。台北。533頁 。
沈世傑。1984b。台灣近海魚類圖鑑。台灣省立博物館。台北。427頁。
沈世傑。1986。中、英、日、拉:世界魚類名典。台灣省立博物館。台北。427頁。
沈世傑、李信徹、邵廣昭、陳哲聰、陳春暉、莫顯蕎等。1993。台灣魚類誌。國立台
灣大學動物學系,台北。960頁。
沈世傑、吳高逸。1995。以鱗魨亞目之腸卷曲型研究其類緣關係。國立台灣大學動物學研
究所動物學刊第六卷第二期。41-56頁。
邵廣昭。1976。台灣海域產沙梭屬魚類系統分類學之研究。國立台灣大學碩士學論文。
133頁。
林沛立。1994。台灣海域產魚參科魚類之系統分類研究。國立台灣海洋大學碩士學位論文。
175頁。
林麗紅。1987。台灣海域產金梭魚科魚類系統分類學之研究。私立中國文化大學海洋研究
所資源組碩士學位論文。116頁。
徐克學。1996。數量分類學。水產出版社。台北。375頁。
陳正平。1996。利用形態及粒線體去氧核醣核酸的12SrRNA序列探討天竺鯛科魚類分
類與類緣關係。國立台灣大學海洋研究所博士論文。192頁。
陳兼善、于名振。1986。台灣脊椎動物誌(上冊)。台灣商務印書館。台北。442頁。
陳鴻鳴。1997。台灣海域鯙科魚類之系統分類研究。國立海洋大學博士學位論文。141頁。
劉振卿。1981。鯔科魚類的生物學研究。國立台灣大學博士學位論文。203頁。
Alock, A., 1894a. ─ Natural history notes from H.M. Indian Marine Survey Steamer
''Investigator'', Commander C.E. Oldham. R. N., commanding Series 2, No. 11. An account of
a recent collection of bathyal fishes from the Bay of Bengal and from the Laccadive Sea.
Journal of the Asiatic Society of Bengal, 63 (pt.2)(2): 115-137, pls 6-7.
Alock, A., 1894b.─Illustrations of the zoology of the Royal Indian Marine Survey Ship
Investigator. Fishes. Part II, pls VIII-XIII.
Anderson, M.E. 1997. Notes on the Genus [Hymenocephalus] Giglioli, 1884 (Teleostei:
Macrouridae) in Southern Africa. Transactions of the Royal Society of South Africa 51:229-
232.
Arai, T., and T. Iwamoto. 1979. A new species of the macrourid
fish genus Coelorinchus from off Tasmania, New Zealand, and
the Falkland Islands. Japanese Journal of Ichthyology 26(3):
238-246.
Arai, T. and P. J. McMillan 1982. A new Macrourid fish, Coelorinchus biclinozalis
from New Zealand, and redescription of C. Australis from Australia. Japan. J.
Ichthyol. 29(2): 115-116.
FAO 1985. Macrouridae (sometimes Coryphaenoididae and
Macrouroididae, in part, in literature), Grenadiers. FAO
Species Identification Sheets :288-301.
Gilbert, C. H. AND F. Cramer.1897. Report on the fishes dredged in deep water near the
Hawaiian Islands, with descriptions and figures of twenty-three new species. Proc. U. S.
Natl. Mus. 19: 403-435.
Gilbert, C. H.1905. The deep-sea fishes of the Hawaiian Islands. Pp575-713, figs. 230-276, pls.
66-101 in The aquatic resources of the Hawaiian Island, D. S. Jordan and B. W. Evemann,
eds. Bull. U. S. Fish Comm. 1903, 22(pt.2, sect.2).
Gilbert, C. H., and C, L. Hubbs, 1916. Report on the Japanese macrouriod fishes collected by the
United States Fisheries Steamer "Albatross" in 1906, with a synopsis of the genera. Proc. U.
S. Nat. Mns. 51 (2149): 135-214, 3 pls.
Gilbert, C. H., and C. L. Hubbs 1920. The Macrouroid fishes of the Philippine islands
and the east indies. United States National Museum. Bull. 100, Vol. 1. Part 7.
Iwamato, Tomio, 1986. Family Macrourida. pp. 330-341 in Smith, M. M. and
P. C. Heemstra, editors, Smith’s Sea Fishes.
Howes, G. J AND O. A. Crimmen. 1990. A review of the Bathygadidae (Teleostei:
Gadiformes). Bull. British Mus. Nat. Hist. (Zool.) 56(2):155-203.
Hubbs, C.L., and T. Iwamoto. 1977. A New Genus ([Mesobius]), and Three New Bathypelagic
Species of Macrouridae (Pisces, Gadiformes) from the Pacific Ocean. Proceedings of the
California Academy of Sciences 41(7):233-251.
Iwamoto, T., and D.L. Stein. 1974. A Systematic Review of the Rattail Fishes (Macrouridae:
Gadiformes) from Oregon and Adjacent Waters. Occasional Papers of the California
Academy of Sciences (111):1-79.
Iwamoto, T. 1979. Eastern pacific macrourine grenadiers with seven branchiostrgal rays (Pisces:
Macrouridae). Proceedings of the California Academy of Sciences 42(5):135-179.
Iwamoto,T.1982. Ventrifossa johnboborum, a new grenadier from the Western Pacific
(Macrouridae: Pisces). Aust.Zool.21(1):55-61.
Iwamoto, T., and T. Arai. 1987. A new grenadier Malacocephalus okamurai (Pisces: Gadiformes:
Macrouridae) from the Western Atlantic. Copeia (1):204-208.
Iwamoto, T., and Y.I. Sazonov. 1988. A review of the southeastern Pacific Coryphaenoides (Sensu
Lato) (Pisces, Gadiformes, Macrouridae). Proceedings of the California Academy of
Sciences 45(3):35-82.
Iwamoto, T. 1989. Phylogeny of grenadiers (suborder Macrouroidei): Anothe. Or
interpretation in Cohen(ed. ). Papers on the systematics of Gadiform Fishes. Sci. Ser. nat.
Hist. Mus. L.A. Co., 32 pp. 159-174.
Iwamoto, T. 1990. Taxonomy of Macrouridae. in FAO species catalogue. Vol. 10 Gadiform fishes
of the world (Order Gadiformes). An annotated and illustrated catalogue of cods, hakes.
grenadiers and other Gadiform fishes know to date. FAO Fisheries Synopsis No. 125, Vol.
10. Rome, FAO. 442 pp.
Iwamoto,T.1994. Revision of the Genus Kumba (Pisces, Gadiformes,Macrouridae), with the
Description of Three New Species. Proceedings of the California Academy of Sciences
48(11): 221-237.
Iwamoto,T.1997. [Trachonurus robinsi], a New Species of Grenadier (Gadiformes, Macrouridae)
fromthe Philippines. Bulletin of Marine Science 60(3):942-949.
Iwamoto, T., and P. McMillan. 1997. A New Grenadier, Genus [Trachonurus], from New Zealand
and Australia (Pisces: Gadiformes: Macrouridae). Memoirs of the Museum of Victoria
56(1):255-259.
Iwamoto, T. and M. Eric Anderson 1999. New species of Caelorinchus(Macrourida, Gadiformes,
Teleostei) from the Indian Ocean. Copeia, 1999(4), 1079-1083 pp.
Iwamoto, T., P. McMillan, and Y. N. Shcherbachev. 1999. A new grenadier, genus Caelorinchus
from Australia and New Zealand (Pisces, Gadiformes, Macrouridae). New Zeal. J. Mar.
Freshw. Res. 33: 49-54.
Iwamoto, T. and A. Williams. 1999. Grenadiers (Pisces, Gadiformes) from the continental slope
of western and northwestern Australia. Proc. Calif. Acad. Sci. 51(3): 105-243.
Katayama, M. 1942. A new Macrouroid fish from the Japan Sea. Zool. Mag. (Tokyo)
54(8): 332-334.
Marshall, N.B. and D.M. Cohen 1973. Order Anacanthini(Gadiformes) — Characters and synopsi of familes. Memoir Sears Foundation for Marine Research, I(6) 479-495.
McCann, C. and D.G. Mcknight 1980. The Marine fauna of new Zealand: Macrourid
fishes(Pisces: Gadia). New Zealand Oecanographic Institute Memoir 61, 90 pp.
McCann, C., and D. G. Mcknight 1980. The marine fauna of New Zealand Macrouid
fishes (Pisces: Gadida). New Zealand Oceanographic Institute Memoir 61.
McMillan, P.J., and C.D. Paulin. 1993. Descriptions of Nine New Species of Rattails of the Genus
[Caelorinchus] (Pisces, Macrouridae) from New Zealand. Copeia (3):819-840
McMillan, P.J. 1999. New grenadier fishes of the genus [Coryphaenoides] (Pisces: Macrouridae);
one from off New Zealand and one widespread in the southern Indo-West Pacific and
Atlantic Oceans. New Zealand Journal of Marine and Freshwater Research 33:481-489.
Merrett, N.R. 1980. A new species of the deep-sea fish genus Coelorinchus Giorna (Macrouridae)
from the eastern North Atlantic, with notes on its ecology. J. Fish Biol. 17():553-
564.
Merrett,N.R.1983. A new species of the deep-sea fish genus Coryphaenoides Gunnerus
(Macrouridae) from the tropical eastern North Atlantic and its relationship with C.
(Coryphaenoides) marshalli Iwamoto,1970. J. Fish Biol. 22(): 265-278.
Merrett,N.R.1986. Macrouridae of the eastern North Atlantic. Merrett, N.R. 1986. Macrouridae of
the eastern North Atlantic. Fich.Ident. ():1-14.
Merrett, N.R., and S.H. Barnes. 1996. Preliminary Survey of Egg Envelope Morphology in the
Macrouridae and the Possible Implications of its Ornamentation. J. Fish Biol. (48):101-
119.
Okamura, O. 1970a. Fauna Japonica, Macrourina (Pisces). Tokyo: Academic Press of Japan,
ii + 216 pp., 44 pls.
Okamura,O. 1970b. Studies on the macrouroid fishes of Japan. Morphology, ecology and
phylogeny. Usa Marine Biological Station Report 17(1-2): 1-179.
Okamura O.Relationships of the Suborder Macrouroidei and Related
Groups with Comments on Merlucciidae and Steindachneria. Natural
History Museum of Los Angeles County Science Series (32). 1989. 129-142.
Sazonov, Y.I., and Y.N. Shcherbachev. 1997. The Composition and Some Problems Pertainng to
the Variatability of Species of Subgenus [Sokodara] (Genus Ventrifossa, Macrouridae).
Japanese Journal of Ichthyology 37(7):529-533.
Shcherbachev, Y.N., and T. Iwamoto. 1995. Indian Ocean Grenadiers of the Subgenus
[Coryphaenoides], Genus [Coryphaenoides] (Macrouridae, Gadiformes, Pisces).
Proceedings of the California Academy of Sciences 48(14): 285-314.
Shcherbachev, Y.N.,Y.I. Sazonov, and T. Iwamoto. 1992. Synopsis of the Grenadier Genus
Kuronezumia (Pisces: Gadiformes: Macrouridae), with Description of a New Species.
():695- 706.
Smith, J.L.B. 1968. New and Interesting Fishes from Deepish Water off Durban, Natal and
Southern Mozambique. Invest. Rep. of The Oceanographic Res. Inst. (19):1-30.
QRCODE
 
 
 
 
 
                                                                                                                                                                                                                                                                                                                                                                                                               
第一頁 上一頁 下一頁 最後一頁 top
系統版面圖檔 系統版面圖檔