臺灣博碩士論文加值系統

早在1717年，諸羅縣志即出現有關台灣山茶的記載。不過，歷來僅有地方人士利用之。直到1999年茶業改良場成功培育出以台灣山茶為父本的優良紅茶品種，它才開始被受到重視。然而，台灣山茶的分類至今仍然模糊不清，使得它的應用及保育基礎相當薄弱。為使未來相關於台灣山茶的研究以及應用能順利進行，解決最基礎的分類問題是當前首要之工作，此即本論文之目標。本論文總共分為七章。第一章先對台灣山茶的定義、基本特性以及相關的分類史做一介紹，以利後續章節相關問題之探討。

第二章乃針對武威山茶與台灣山茶在分類上的關係做一釐清。因為長久以來缺乏活體與標本可供研究，所以武威山茶一度被認為與台灣山茶乃同一分類群。直到2004年元月武威山茶重新被發現，才得以證明兩者確為不同植物。尤有甚者，武威山茶之形態特徵顯示其應隸屬於烏皮茶屬，因此本人將之重新進行分類處理，稱為武威山烏皮茶。本章的研究成果已經發表在Taiwania (2004, 49: 201-208).

第三章則使用外部形態特徵，以數值分類方法來研究台灣山茶以及其兩個近緣分類群—茶與阿薩姆茶的關係。聚類分析與非線性主成份分析之結果，皆支持台灣山茶為相對獨立的一個分類群。而台灣山茶與另兩者最大之不同，在於台灣山茶之新芽為光滑或僅具稀疏之毛，而另兩者皆密被毛；此外台灣山茶的子房光滑，而另兩者則密被毛。其它如嫩枝、葉柄以及葉下表面之毛被狀況，亦可做為分群之輔助。在營養特徵的分析中，台灣山茶的台東族群，呈現與西部的族群較大之差異，暗示分布於台東者為比較特殊之一群。本章的研究成果已經被Taiwania接受。

第四章則是探討台灣山茶、茶與阿薩姆茶在花粉形態上的差異。結果發現，台灣山茶在花粉外壁內層的雕紋與花粉極面的形狀上，與另兩者有所不同，此點再度說明台灣山茶的確為不同於茶與阿薩姆茶的植物。至於不同地區的台灣山茶之間、或是茶與阿薩姆茶之間，皆無法以花粉形態來區分。

第五章乃針對台灣山茶、茶以及阿薩姆茶在氣孔形態及葉片橫切面的解剖構造進行比較。在氣孔形態方面，氣孔周圍角質層突起與異型氣孔的具備與否，可以將所有的樣本分成兩群。在台灣山茶中，除南投的族群之外，皆屬不具有窄肋狀角質層突起與異型氣孔之類群，然而南投族群卻和茶以及阿薩姆茶一樣，皆具有窄肋狀角質層突起與異型氣孔。因此，氣孔形態支持南投族群與其它族群間存在著差異。至於葉片解剖部分，則無顯著的特徵差異可供分群之用。但是根據初步之定性觀察，台灣山茶之葉部內含石細胞密度甚高，此點或可能在未來以統計方法取樣及分析，為台灣山茶之分類提供更多之證據。

第六章嚐試以DNA定序法，來探討台灣山茶與兩近緣分類群的關係。以最大簡約法及鄰近連接法對RPB2 intron 11-16與intron 23兩段序列進行親緣分析，並計算類群間的平均遺傳距離，用以比較各類群在DNA上的差距。結果支持台灣山茶為單源群，且與茶和阿薩姆茶關係較為疏遠。在台灣山茶之中，採自南投與台東的樣本較為接近，而嘉義至屏東的樣本則獨立成另一群。此外，台灣山茶與茶或是阿薩姆茶間的遺傳距離，皆明顯大於茶與阿薩姆茶間的遺傳距離。

第七章則根據前面的研究結果，對台灣山茶進行分類處理。由於台灣山茶相對於茶或是阿薩姆茶皆有較大之差異，基於分類學家對茶與阿薩姆茶所進行的分類處理，本人將台灣山茶處理成種之階級，學名稱為Camellia formosensis (Masamune et Suzuki) M. H. Su, C. F. Hsieh et C. H. Tsou。同時，因為台灣山茶之台東族群在外部形態上已有明顯分化，所以將之處理成變種並命名為C. formosensis var. yungkangensis M. H. Su, C. F. Hsieh et C. H. Tsou，中名稱之為永康山茶。

The Taiwan native wild tea plant was first documented in 1717 on Chu-lo-hsien-chih. The utilization of this plant is mostly restricted to local people until 1999, when a black tea clone was bred by the Tea Research and Extension Station. The researches, applications and conservations on the Taiwan native wild tea plant were limited due to several uncertainty about its taxonomy. This dissertation is set out to elucidate the taxonomic status of the Taiwan native wild tea plant based on morphological and molecular evidences.

There are seven chapters in the dissertation. In chapter 1, the definition, background and taxonomic history of the Taiwan native wild tea plant were introduced.

Chapter 2 concerns about the Camellia buisanensis and the Taiwan native wild tea plant. Because of a lack of materials for examination, some taxonomists thought C. buisanensis is the same with the Taiwan native wild tea plant until living plants were rediscovered in January 2004. Differences between the two taxa become apparent. Morphological data indicate that C. buisanensis is indeed a member of the genus Pyrenaria. Therefore, C. buisanensis was treated as Pyrenaria buisanensis, the results were published in Taiwania (2004, 49: 201-208).

In Chapter 3, the relationship between the Taiwan native wild tea plant and its two closely related taxa - C. sinensis var. sinensis and C. sinensis var. assamica - was explored by numerical methods using morphological characters. Results of the cluster analysis and nonlinear principal component analysis supported the Taiwan native wild tea plant as a distinct taxon. Buds of the Taiwan native wild tea plant are glabrous or sparsely hairy, while those of C. sinensis var. sinensis and C. sinensis var. assamica are densely hairy. Ovaries of the Taiwan native wild tea plant are glabrous, while those of C. sinensis var. sinensis and C. sinensis var. assamica are densely hairy. Vegetative characteristics of the Taiwan native wild tea plant from Taitung are different from individuals in other area, suggesting differentiation or specialization in the Taitung population. The results had been accepted by Taiwania.

In Chapter 4, the morphology of pollens from the Taiwan native wild tea plant, C. sinensis var. sinensis and C. sinensis var. assamica were compared. The pollens of Taiwan native wild tea plant were different from the other two closely related taxa in the sculpture of nexine and the shape at the polar side. In contrast, C. sinensis var. sinensis and C. sinensis var. assamica couldn''t be distinguished by the morphology of pollens.

In Chapter 5, the stomatal and leaf anatomy of the Taiwan native wild tea plant, C. sinensis var. sinensis and C. sinensis var. assamica were compared. As to the stomatal morphology, the Taiwan native wild tea plant except for the Nantou population didn''t have the ribbed cuticles and heteromorphic stomata, while Nantou population, C. sinensis var. sinensis and C. sinensis var. assamica did. Therefore, stomatal morphology supported Nantou population is special. There was no markly difference in leaf anatomy. The density of stone cells in leaves of the Taiwan native wild tea plant seems to be higher than the other two taxa. However, future studies to quantify this character might provide further evidence on taxonomic properties.

In Chapter 6, phylogenetic analyses were conducted using sequences of two DNA fragments, RPB2 intron 11-16 and intron 23. Results by the maximum parsimony and neighbor joining supported the Taiwan native wild tea plant is monophyletic. The genetic distance between the Taiwan native wild tea plant and C. sinensis var. sinensis, or C. sinensis var. assamica, was larger than that between C. sinensis var. sinensis and C. sinensis var. assamica. Within the Taiwan native wild tea plant, Nantou and Taitung populations were similar than Chiayi-Pingtung population.

In Chapter 7, taxonomic treatment on the Taiwan native wild tea plant was proposed, according to results of the previous chapters. Because the Taiwan native wild tea plant is distinct from C. sinensis var. sinensis and C. sinensis var. assamica, a specific rank for the Taiwan native wild tea plant was proposed. The name was given as Camellia formosensis (Masamune et Suzuki) M. H. Su, C. F. Hsieh, et C. H. Tsou. Meanwhile, the Taitung population was named as C. formosensis var. yungkangensis M. H. Su, C. F. Hsieh, et C. H. Tsou, for its markedly differences in the vegetative morphology.

口試委員會審定書 i
誌謝 ii
中文摘要 iii
英文摘要 v
第一章、緒論 1
第一節、台灣山茶名稱之定義 1
第二節、古籍中有關台灣山茶之記載 1
第三節、台灣山茶之分類回顧 1
第四節、台灣山茶之地理分布 5
第五節、台灣山茶之經濟用途 6
第六節、研究目的 7
第二章、台灣山茶與武威山茶之關係 9
第一節、本章緒言 9
第二節、武威山茶之再發現始末 9
第三節、武威山茶之分類處理 11
第四節、武威山烏皮茶之現況及保育 13
第五節、本章結語 13
第三章、台灣山茶與近緣分類群之形態比較 19
第一節、本章緒言 19
第二節、研究方法 19
第三節、結果 24
第四節、討論 30
第五節、本章結語 34
第四章、台灣山茶與近緣分類群之花粉形態比較 35
第一節、本章緒言 35
第二節、研究方法 36
第三節、結果 37
第四節、討論 38
第五節、本章結語 40
第五章、台灣山茶與近緣分類群之葉部解剖比較 47
第一節、本章緒言 47
第二節、研究方法 48
第三節、結果 50
第四節、討論 52
第五節、本章結語 54
第六章、台灣山茶與近緣分類群之DNA序列比較 61
第一節、本章緒言 61
第二節、研究方法 64
第三節、結果 67
第四節、討論 75
第五節、本章結語 77
第七章、台灣山茶之分類處理 79
第一節、綜合回顧與分類處理思考 79
第二節、分類處理 81
參考文獻 91
附錄一、營養特徵原始資料矩陣 99
附錄二、花部特徵原始資料矩陣 105
附錄三、RPB2 intron 11-16原始資料矩陣 107
附錄四、RPB2 intron 23原始資料矩陣 117

Angiosperm Phylogeny Group. 2003. An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG II. Bot. J. Linn. Soc. 141: 399-436.
Ao, C.-Q., G.-X. Chen, G.-P. Zhang, K. Gao, and H.-T. Chang. 2002. Study on micromorphology of pollen exine surface in Camellia. Acta Bot. Yunnanica 24: 619-626.
Bailey, C. D., T. G. Carr, S. A. Harris, and C. E. Hughes. 2003. Characterization of angiosperm nrDNA polymorphism, paralogy, and pseudogenes. Mol. Phylogenet. Evol. 29: 435-455.
Balasaravanan, T., P. K. Pius, R. Raj Kumar, N. Muraleedharan, and A. K. Shasany. 2003. Genetic diversity among south Indian tea germplasm (Camellia sinensis, C. assamica and C. assamica spp. lasiocalyx) using AFLP markers. Plant Science 165: 365-372.
Banerjee, B. 1992a. Botanical classification of tea. In: Willson, K. C. and M. N. Clifford (eds.), Tea: cultivation to consumption. Chapman & Hall, London, U. K. pp. 25-51.
Banerjee, B. 1992b. Selection and breeding of tea. In: Willson, K. C. and M. N. Clifford (eds.), Tea: cultivation to consumption. Chapman & Hall, London, U. K. pp. 53-86.
Bayly, M. J., A. V. Kellow, P. J. D. Lange, K. A. Mitchell, K. R. Markham, P. J. Garnock-Jones, and P. J. Brownsey. 2003. Geographic variation in morphology and flavonoid chemistry in Hebe pubescens and H. bollonsii (Scrophulariaceae), including a new infraspecific classification for H. pubescens. N. Z. J. Bot. 41: 23-53.
Binns, S. E., B. R. Baum, and J. T. Arnason. 2002. A taxonomic revision of Echinacea (Asteraceae: Heliantheae). Syst. Bot. 27: 610-632.
Bland, J. M. and D. G. Altman. 1997. Statistics notes: Cronbach''s alpha. Brit. Med. J. 314: 572.
Chang, H.-T. and S.-X. Ren. 1991. New species of Theaceae from Tropical Asia. Acta Sci. Nat. Univ. Sunyatseni 30: 67-71.
Chen, J., P. Wang, Y. Xia, M. Xu, and S. Pei. 2005. Genetic diversity and differentiation of Camellia sinensis L. (cultivated tea) and its wild relatives in Yunnan province of China, revealed by morphology, biochemistry and allozyme studies. Genet. Resour. Crop Evol. 52: 41-52.
Chen, L. and S. Yamaguchi. 2002. Genetic diversity and phylogeny of tea plant (Camellia sinensis) and its related species and varieties in the section Thea genus Camellia determined by randomly amplified polymorphic DNA analysis. J. Hortic. Sci. Biotech. 77: 729-732.
Chen, L. and S. Yamaguchi. 2005. RAPD markers for discriminating tea germplasms at the inter-specific level in China. Plant Breed. 124: 404-409.
Cheng, W.-T. 1990. Populational studies on Formosan Lilium (Liliaceae) I. A cluster analysis of variants in L. longiflorum Thunb. Taiwania 35: 198-205.
Cheng, Y.-P., S.-Y. Hwang, and T.-P. Lin. 2005. Potential refugia in Taiwan revealed by the phylogeographical study of Castanopsis carlesii Hayata (Fagaceae). Mol. Ecol. 14: 2075-2085.
Crawford, D. J. 2000. Plant macromolecular systematics in the past 50 years: one view. Taxon 49: 479-501.
Cronbach, L. J. 1951. Coefficient alpha and the internal structure of tests. Psychometrika 16: 297-334.
Cronquist, A. 1981. An integrated system of classification of flowering plants. Columbia University Press, New York. USA. 1262pp.
de Leeuw, J. 1982. Nonlinear principal components analysis. In: Caussinus, H., P. Ettinger, and R. Tomassone (eds.), COMPSTAT Proceedings in Computational Statistics, Physica Verlag, Vienna, Austria. pp. 77-89.
Denton, A. L., B. L. Mcconaughy, and B. D. Hall. 1998. Usefulness of RNA polymerase II coding sequences for estimation of green plant phylogeny. Mol. Biol. Evol. 15: 1082-1085.
Devarumath, R. M., S. Nandy, V. Rani, S. Marimuthu, N. Muraleedharan, and S. N. Raina. 2002. RAPD, ISSR and RFLP fingerprints as useful markers to evaluate genetic integrity of micropropagated plants of three diploid and triploid elite tea clones representing Camellia sinensis (China type) and C. assamica ssp. assamica (Assam-India type). Plant Cell Rep. 21: 166-173.
Dunn, G. and B. S. Everitt. 1982. An introduction to mathematical taxonomy. Cambridge University Press, London, U. K. 152pp.
Ellis, R. N., P. M. Kroonenberg, B. D. Harch, and K. E. Basford. 2006. Non-linear principal components analysis: an alternative method for finding patterns in environmental data. Environmetrics 17: 1-11.
Erdtman, G. 1952. Pollen morphology and plant taxanomy: Angiosperms. Waltham, Mass, Stockholm, Sweden. 539 pp.
Freeman, S., J. West, C. James, V. Lea, and S. Mayes. 2004. Isolation and characterization of highly polymorphic microsatellites in tea (Camellia sinensis). Mol. Ecol. Notes 4: 324-326.
Gower, J. C. 1971. A general coefficient of similarity and some of its properties. Biometrics 27: 857-871.
Greuter, W., J. Mcneill, F. R. Barrie, H. M. Burdet, V. Demoulin, T. S. Filgueiras, D. H. Nicolson, P. C. Silva, J. E. Skog, P. Trehane, N. J. Turland, and D. L. Hawksworth (eds.). 2000. International Code of Botanical Nomenclature (Saint Louis Code). Adopted by the Sixteenth International Botanical Congress St. Louis, Missouri, July–August 1999 (Regnum veg. 138). Koeltz Scientific Books, Konigstein, Germany.
Gugerli, F. 1997. Hybridization of Saxifraga oppositifolia and S. biflora (Saxifragaceae) in a mixed alpine population. Plant Syst. Evol. 207: 255-272.
Henry, A. 1896. A list of plants from Formosa. Trans. Asiat. Soc. Jap. 4. Sup. p. 20.
Hickey, L. J. and J. A. Doyle. 1977. Early Cretaceous fossil evidence for angiosperm evolution. Bot. Rev. 43: 3-104.
Hsieh, C.-F., L.-K. Ling, and K.-C. Yang. 1996. Theaceae. In: Huang, T.-C. et al. (eds.), Flora of Taiwan, 2nd ed. 2: 662-693. Editorial Committee, Dept. Bot., NTU, Taipei, Taiwan.
Huang, T.-C. 1972. Pollen Flora of Taiwan. Ching-hwa Press Co., LTD., Taipei, Taiwan. 297pp.
Hutchinson, J. 1973. The families of flowering plants. The Clarendon Press, Oxford, UK. 968pp.
IUCN. 2001. IUCN red list categories and criteria: Version 3.1. IUCN Species Survival Commission. IUCN, Gland, Switzerland and Cambridge, UK. pp. 14-15.
Jolliffe, I. T. 1986. Principal component analysis Springer-Verlag, New York, USA. 271pp.
Jones Jr., S. B. and A. E. Luchsinger. 1987. Plant systematics. McGraw-Hill. USA. p. 4.
Judd, W. S. , C. S. Campbell, E. A. Kellogg and P. F. Stevens. 2002. Plant systematics: a phylogenetic approach. Sinauer Associates, Inc. Sunderland, U.S.A. p. 33-37.
Katoh, Y., M. Katoh, Y. Takeda, and M. Omori. 2003. Genetic diversity within cultivated teas based on nucleotide sequence comparison of ribosomal RNA maturase in chloroplast DNA. Euphytica 134: 287-295.
Kaundun, S. S., A. Zhyvoloup, and Y.-G. Park. 2000. Evaluation of the genetic diversity among elite tea (Camellia sinensis var. sinensis) accessions using RAPD markers. Euphytica 115: 7-16.
Keng, H. 1950. The Theaceae of Taiwan. Taiwania 1: 1-233.
Keng, H. 1962. Comparative morphological studies in Theaceae. Univ. Calif. Publ. Bot. 33: 269-384.
Keng, H. 1973. Nomenclatural note on Wu Pi Cha or the black-barked tea plant. Taiwania 18: 55-56.
Lai, J.-A., W.-C. Yang, and J.-Y. Hsiao. 2001. An assessment of genetic relationships in cultivated tea clones and native wild tea in Taiwan using RAPD and ISSR markers. Bot. Bull. Acad. Sin. 42: 93-100.
Larkin, R. and T. Guilfoyle. 1993. The second largest subunit of RNA polymerase II from Arabidopsis thaliana. Nucleic Acids Res. 21: 1038.
Li, H.-L. 1976. Theaceae. In: Li, H.-L.et al. (eds.), Flora of Taiwan. Vol. 2. Epoch Publ. Co., Taipei, Taiwan. pp. 592-619.
Liu, T.-S. and F.-Y. Lu. 1967. Studies in the Taiwan Theaceae based on the morphological characters of leaves. Tech. Bull. Exp. For., Nat. Taiwan Univ. 52: 1-55.
Luna, I. and H. Ochoterena. 2004. Phylogenetic relationships of the genera of Theaceae based on morphology. Cladistics 20: 223-270.
Matheny, P. B. 2005. Improving phylogenetic inference of mushrooms with RPB1 and RPB2 nucleotide sequences (Inocybe; Agaricales). Mol. Phylogenet. Evol. 35: 1-20.
Mckenzie, R. J., J. M. Ward, J. D. Lovis, and I. Breitwieser. 2004. Morphological evidence for natural intergeneric hybridization in the New Zealand Gnaphalieae (Compositae): Anaphalioides bellidioides × Ewartia sinclairii. Bot. J. Linn. Soc. 145: 59-75.
Merrill, E. D. and F. P. Metcalf. 1937. New Kwangtung plants. Lingnan Sci. J. 16: 172.
Mondal, T. K. 2002. Assessment of genetic diversity of tea (Camellia sinensis (L.) O. Kuntze) by inter-simple sequence repeat polymerase chain reaction. Euphytica 128: 307-315.
Mondal, T. K., A. Bhattacharya, M. Laxmikumaran, and P. S. Ahuja. 2004. Recent advances of tea (Camellia sinensis) biotechnology. Plant Cell, Tiss. Organ Cult. 76: 195-254.
Muvaffak, A., M. Dogan, and C. C. Bilgin. 2001. A numerical taxonomic study of the genus Acantholimon Boiss. (Plumbaginaceae) in Ankara Province (Turkey). Isr. J. Plant Sci. 49: 298-300.
Nickerson, J. and G. Drouin. 2004. The sequence of the largest subunit of RNA polymerase II is a useful marker for inferring seed plant phylogeny. Mol. Phylogenet. Evol. 31: 403-415.
Nishimoto, Y., O. Ohnishi, and M. Hasegawa. 2003. Topological incongruence between nuclear and chloroplast DNA trees suggesting hybridization in the urophyllum group of the genus Fagopyrum (Polygonaceae). Genes Genet. Syst. 78: 139-153.
Oshida, T., J.-K. Lee, L.-K. Lin, and Y.-J. Chen. 2006. Phylogeography of Pallas''s squirrel in Taiwan: geographical isolation in an arboreal small mammal. J. Mammal. 87:247-254.
Oxelman, B. and B. Bremer. 2000. Discovery of paralogous nuclear gene sequences coding for the second-largest subunit of RNA Polymerase II (RPB2) and their phylogenetic utility in Gentianales of the asterids. Mol. Biol. Evol. 17: 1131-1145.
Oxelman, B., N. Yoshikawa, B. L. Mcconaughy, J. Luo, A. L. Denton, and B. D. Hall. 2004. RPB2 gene phylogeny in flowering plants, with particular emphasis on asterids. Mol. Phylogenet. Evol. 32: 462-479.
Paul, S., F. N. Wachira, W. Powell, and R. Waugh. 1997. Diversity and genetic differentiation among populations of Indian and Kenyan tea ( Camellia sinensis (L.) O. Kuntze) revealed by AFLP markers. Theor. Appl. Genet. 94: 255-263.
Pearson, D. L. 1994. Selecting indicator taxa for the quantitative assessment of biodiversity. Phil. Trans. R. Soc. Lond. B. 345: 75-79.
Popp, M. and B. Oxelman. 2001. Inferring the history of the polyploid Silene aegaea (Caryophyllaceae) using plastid and homoeologous nuclear DNA sequences. Mol. Phylogenet. Evol. 20: 474-481.
Popp, M. and B. Oxelman. 2004. Evolution of a RNA polymerase gene family in Silene (Caryophyllaceae)—incomplete concerted evolution and topological congruence among paralogues. Syst. Biol. 53: 914-932.
Prabhakar, M. 2004. Structure, delimitation, nomenclature and classification of stomata. Acta Bot. Sin. 46: 242-252.
Prince, L. M. and C. R. Parks. 2001. Phylogenetic relationships of Theaceae inferred from chloroplast DNA sequence data. Am. J. Bot. 88: 2309-2320.
Roncal, J., J. Francisco-Ortega, C. B. Asmussen, and C. E. Lewis. 2005. Molecular phylogenetics of tribe Geonomeae (Arecaceae) using nuclear DNA sequences of phosphoribulokinase and RNA polymerase II. Syst. Bot. 30: 275-283.
Sasaki, S. 1931. Miscellaneous contributions to the flora of Formosa (X). Trans. Nat. His. Soc. Form. 21: 222.
Schultze-Motel, J. and D. Meyer. 1981. Numerical taxonomic studies in the genera Triticum L. and Pisum L. Genet. Resour. Crop Evol. 29: 241-250.
Sealy, J. 1958. A revision of the genus Camellia. Royal Horticultural Society, London, U. K. p. 119.
Shaw, J., E. B. Lickey, J. T. Beck, S. B. Farmer, W. Liu, J. Miller, K. C. Siripun, C. T. Winder, E. E. Schilling, and R. L. Small. 2005. The tortoise and the hare II: relative utility of 21 noncoding chloroplast DNA sequences for phylogenetic analysis. Am. J. Bot. 91: 142-166.
Sneath, P. H. A. 1995. Thirty years of numerical taxonomy. Syst. Biol. 44: 281-298.
Sneath, P. H. A. and R. R. Sokal. 1973. Numerical taxonomy: the principles and practice of numerical classification. W. H. Freeman and Company, San Francisco, USA. 573pp.
Sokal, R. R. 1986. Phenetic taxonomy: theory and methods. Annu. Rev. Ecol. Syst. 17: 423-442.
Soltis, D. E. and P. S. Soltis. 1998. Choosing an approach and an appropriate gene for phylogenetic analysis. In D. E. Soltis, P. S. Soltis and J. J. Doyle (eds.), Molecular systematics of plants II: DNA sequencing. Kluwer Academic Publishers, Boston, USA. pp. 1-42.
Stefanovic, S. and R. G. Olmstead. 2004. Testing the phylogenetic position of a parasitic plant (Cuscuta, Convolvulaceae, Asteridae): Bayesian inference and the parametric bootstrap on data drawn from three genomes. Syst. Biol. 53: 384-399.
St-Laurent, L., B. R. Baum, K. Akpagana, and J. T. Arnason. 2000. A numerical taxonomic study of Trema (Ulmaceae) from Togo, West Africa. Syst. Bot. 25: 399-413.
Stuessy, T. F. 1990. Plant taxonomy: the systematic evaluation of comparative Data. Columbia University Press, New York, USA. 514pp.
Su, H.-J. 1984. Studies on the climate and vegetation types of the natural forests in Taiwan (II): altitudinal vegetation zone in relation to temperature gradient. Quart. J. Chin. For. 17: 57-73.
Su, M.-H., S.-Z. Yang, and C.-F. Hsieh. 2004. The identity of Camellia buisanensis Sasaki (Theaceae). Taiwania 49: 201-208.
Swofford, D. L. 2001. PAUP* . Phylogenetic analysis using parsimony (*and other methods). Version 4. in. Sinauer Associates, Sunderland, Massachusetts.
Takeda, Y. 1990. Cross compatibility of tea (Camellia sinensis) and its allied species in the genus Camellia. JARQ 24: 111-116.
Takeda, Y. 1994. Differences in caffeine and tannin contents between tea cultivars, and application to tea breeding. JARQ 28: 117-123.
Takeo, T. 1992. Green and semi-fermented teas. In Willson, K. C. and M. N. Clifford (eds.), Tea: cultivation to consumption, 1 Ed., Vol. Chapman & Hall, London, U. K. pp. 413-457.
Tang, S. and Y. Zhong. 2002. A phylogenetic analysis of nrDNA ITS sequences from Ser. Chrysantha (Sect. Chrysantha, Camellia, Theaceae). J. Genet. Mol. Biol. 13: 105-107.
Thompson, J. D., T. J. Gibson, F. Plewniak, F. Jeanmougin, and D. G. Higgins. 1997. The CLUSTAL_X windows interface: flexible strategies for multiple sequence alignment aided by quality analysis tools. Nucleic Acids Res. 25: 4876-4882.
Tsou, C.-H. 1997. Embryology of the Theaceae - anther and ovule development of Camellia, Franklinia, and Schima. Am. J. Bot. 84: 369-381.
Tsou, C.-H. 1998. Early floral development of Camellioideae (Theaceae). Amer. J. Bot. 85: 1531-1547.
Wachira, F. N., W. Powell, and R. Waugh. 1997. An assessment of genetic diversity among Camellia sinensis L. (cultivated tea) and its wild relatives based on randomly amplified polymorphic DNA and organelle-specific STS. Heredity 78: 603-611.
Wachira, F., J. Tanaka, and Y. Takeda. 2001. Genetic variation and differentiation in tea (Camellia sinensis) germplasm revealed by RAPD and AFLP variation. J. Hortic. Sci. Biotech. 76: 557-563.
Walker, J. W. 1974. Aperture evolution in the pollen of primitive angiosperms. Am. J. Bot. 61: 1112-1137.
Ward, J. M. 1993. Systematics of New Zealand Inuleae (Compositae - Asteraceae) - 2. A numerical phenetic study of Raoulia in relation to allied genera. N. Z. J. Bot. 31: 29-42.
Wei, Z.-X., M. S. Zavada, and T.-L. Ming. 1992. Pollen morphology of Camellia (Theaceae) and its taxonomic significance. Acta Bot. Yunnanica 14: 275-282.
Wolfe, K. H., W.-H. Li, and P. M. Sharp. 1987. Rates of nucleotide substitution vary greatly among plant mitochondrial, chloroplast, and nuclear DNAs. Proc. Natl. Acad. Sci. U. S. A. 84: 9054-9058.
Xiao, T. 2001. Phylogenetic relationships of the genus Camellia (Theaceae) based on the RNA polymerase II (RPB2) sequences. Doctor Thesis, Department of Biology, University of North Carolina. Chapel Hill, USA. 293pp.
Yang, S. X., J. B. Yang, L. G. Lei, D. Z. Li, H. Yoshino, and T. Ikeda. 2004. Reassessing the relationships between Gordonia and Polyspora (Theaceae) based on the combined analyses of molecular data from the nuclear, plastid and mitochondrial genomes. Plant Syst. Evol. 248: 45-55.
Zaharof, E. 1988. A phenetic study of Fritillaria (Liliaceae) in Greece. Plant Syst. Evol. 161: 23-34.
Zavada, M. S. and Z.-X. Wei. 1993. A contribution to the pollen morphology of Camellia. Grana 32: 233-242.
山本由松與森邦彥. 1934. 台灣產山茶科植物ソ種類ソ形態學的鑑識ズ就サ. Ёルビヤ 5: 24-48.
王任翔, 胡長華, 梁盛業與黃松殿. 2002. 金花茶組植物葉表皮特徵的掃描電鏡觀察. 廣西林業科學 31: 133-136.
王任翔, 梁盛業, 李潔榮, 高成偉, 胡長華與張杏輝. 2003. 金花茶組植物葉表皮特徵的掃描電鏡觀察 (II). 廣西林業科學 32: 12-16.
王兩全, 何信鳳, 陳右人, 馮鑑淮與邱再發. 1990. 台灣野生茶樹種原保存及利用 I. 台灣眉原山野生茶樹調查. 台灣茶業研究彙報 9: 1-6.
王奎玲, 黃鑫, 劉慶華, 牟少華, 劉慶超與張啟翔. 2006. 耐冬山茶葉組織結構與品種分類研究. 河南農業大學學報 40: 291-296.
北村四郎. 1950. 茶ごШバワ. 植物分類地理 14: 56-63.
史樨、陳永盛、楊守國、石振源與廖增祿. 1972. 台灣南投野生茶樹之調查. 台灣農業 8：193-207.
正宗嚴敬. 1936. 最新台灣植物總目錄. Kudo編輯部, 台北, 台灣. 第140頁.
朱仕玠. 1765. 小琉球漫誌.（重印本：台灣銀行經濟研究室. 1957. 小琉球漫誌. 台灣文獻叢刊第3種. 台灣銀行. 台灣. 第54頁）.
吳振鐸. 1963. 茶樹花部形態的研究. 中華農學會報 新(44): 34-52.
吳振鐸. 1968. 茶樹葉部解剖及其與茶葉品質的關係研究. 中華農學會報 新(64): 29-43.
吳振鐸, 馮鑑淮與蔡俊明. 1965. 茶樹葉部石核細胞的形態與分佈及其與茶葉品質的關係研究. 中華農學會報 新(52): 42-72.
吳振鐸、賈雲翔、馮鑑淮與蔡俊明. 1970. 台灣眉原山野生茶樹形態之觀察（I）. 台灣農業季刊 6: 1-13.
吳振鐸、馮鑑淮與蔡俊明. 1972. 台灣眉原山野生茶樹形態之觀察（II）. 台灣農業季刊8: 133-160.
呂勝由與楊遠波. 1987. 台灣的野生茶樹—武威山茶. 中華林學季刊 20: 101-107.
呂福原、歐辰雄與呂金誠. 2000. 台灣樹木解說第四冊. 行政院農業委員會. 台北, 台灣. 第12頁.
束際林, 陳亮, 王海思, 王平盛, 許久與宋維希. 1998. 茶樹及其他山茶屬植物花粉形態, 超微結構及演化. 茶葉科學 18: 6-15.
李台強與張清寬. 2003. 台灣茶樹種原圖誌. 行政院農業委員會茶業改良場, 桃園, 台灣. 第173頁.
李廣清, 孫立與劉燕. 2005. 山茶屬連蕊茶組6種植物花粉形態特徵研究. 熱帶亞熱帶植物學報 13: 40-44.
沈中桴. 1997. 台灣的生物地理：2. 一些初步思考與研究. 台灣博物館年刊. 40: 361-450.
周鍾瑄. 1717. 諸羅縣志.（重印本：台灣銀行經濟研究室. 1962. 諸羅縣志. 台灣文獻叢刊第141種. 台灣銀行. 台北, 台灣. 第286頁）.
邱垂豐、邱瑞騰與林金池. 2004. 台茶18號 (別名「紅玉」) 簡介. 茶訊 818: 5.
金平亮三. 1936. 台灣樹木誌. 台灣總督府中央研究所. 第456頁.
胡智益. 2004. 台灣茶樹種原葉部性狀及DNA 序列變異之探討. 國立台灣大學農藝研究所碩士論文. 台北, 台灣. 全146頁.
郁永河. 1697. 裨海紀遊.（重印本：台灣銀行經濟研究室. 1959. 台陽見聞錄. 台灣文獻叢刊第44種. 台灣銀行. 台北, 台灣. 全72頁）.
韋仲新. 1997. 山茶科花粉超微結構及其系統學意義. 雲南植物研究 19: 143-153.
唐紹清, 鐘揚, 施蘇華與張宏達. 2001. Camellia nitidissima與C. petelotii之間的關係研究 - 來自nrDNA ITS的證據. 武漢植物學研究 19: 449-456.
唐贊袞. 1891. 台陽見聞錄.（重印本：台灣銀行經濟研究室. 1958. 台陽見聞錄. 台灣文獻叢刊第30種. 台灣銀行. 台北, 台灣. 第157頁）.
徐英祥. 1995. 台灣日據時期茶業文獻譯集. 台灣省茶業改良場. 第115-122頁.
張宏達. 1981. 山茶屬植物的系統研究. 中山大學學報自然科學版 20: 87-99.
張宏達. 1984. 茶植物資源的修訂. 中山大學學報自然科學版 23: 1-12.
張宏達與任善湘. 1998. 中國植物志第四十九卷第三分冊. 科學出版社. 北京. 中國. 第133-135頁.
張浩與莊雪影. 2004. 山茶科部分屬種葉表皮形態學研究. 華南農業大學學報 25: 87-93.
敖成齊, 陳功錫, 席嘉賓, 鄭政偉與張宏達. 2001. 山茶屬短柱茶組花粉形態的研究. 華南農業大學學報 22: 66-68.
敖成齊, 陳功錫與張宏達. 2002. 山茶屬的葉表皮形態及其分類學意義. 雲南植物研究 24: 68-74.
陳亮, 童啟慶, 高其康, 束際林與虞富蓮. 1997. 山茶屬8種1變種花粉形態比較. 茶葉科學 17: 183-188.
閔天祿. 2000. 世界山茶屬的研究. 雲南科技出版社. 昆明, 中國. 第130-133頁.
黃叔璥. 1724. 台海使槎錄.（重印本：台灣銀行經濟研究室. 1957. 台海使槎錄. 台灣文獻叢刊第4種. 台灣銀行. 台北, 台灣. 第62頁）.
楊世雄. 2000. 廣義核果茶屬的分類、分化和地理分布. 中國科學院昆明植物研究所博士論文. 昆明, 中國. 全201頁.
楊世雄與閔天祿. 1995. 山花科核果茶屬, 石筆木屬和擬核果茶屬的分類學位置. 雲南植物研究 17: 192-196.
楊世雄, 劉愛忠, 彭華與吳征鎰. 2003. 核果茶屬的氣孔器類型及其系統學意義. 廣西植物 23: 250-252.
楊遠波、劉和義與呂勝由. 1997. 台灣維管束植物簡誌第二卷. 行政院農業委員會.台北, 台灣. 第189頁.
鄒天才與婁義龍. 1995. 山茶屬三種植物葉片解剖特徵研究. 貴州科學 13: 11-17.
鄒天才, 張著林, 周洪英與婁義龍. 1996. 山茶屬五種植物葉片解剖特徵及與光合生理相關研究. 西北植物學報 16: 42-51.
鈴木重良. 1937. 台灣植物便覽. 台北, 台灣. 第262頁.
劉棠瑞. 1962. 台灣木本植物圖誌. 國立台灣大學農業學院. 台北, 台灣. 第235頁.
劉業經、呂福原與歐辰雄. 1988. 台灣樹木誌. 國立中興大學農學院. 台中. 台灣. 第491頁.
蔡依真. 2003. 台灣茶樹種源遺傳歧異度研究與分子指紋之建立. 國立台灣大學農藝研究所碩士論文. 台北, 台灣. 全81頁.
蔡淑華. 1972. 植物解剖學. 國立編譯館, 台北, 台灣. 第37-46頁.
鄧六中, 黃紫恆與漆龍霖. 1992. 山茶屬主要油用物種葉片解剖特徵的研究. 經濟林研究 10: 8-15.
橋本實. 1967. 台灣產野生茶ズ關する研究. 日本名城大學農學部學術報告 4: 17-20.
應紹舜. 1995. 台灣產茶科植物分類之研究. 台大農學院研究報告 35(2): 131-170.
戴素賢與謝赤軍. 2001. 七種高香型烏龍茶葉片結構的比較. 廣東茶業: 28-29.
藍鼎元. 1732. 東征集.（重印本：台灣銀行經濟研究室. 1958. 東征集. 台灣文獻叢刊第12種. 台灣銀行. 台北, 台灣. 第85頁）.
嚴學成. 1982. 茶葉的表皮細胞. 植物學報 24: 312-318.
蘇夢淮. 1993. 南仁山亞熱帶雨林樹冠層葉片結構之研究. 國立台灣大學植物學研究所碩士論文. 台北, 台灣.