布氏鯨(Balaenoptera edeni)屬於鬚鯨科，廣泛分布於熱帶及亞熱帶海域，牠們沒有長距離的洄游習性。出現於土佐灣的布氏鯨族群在親緣上屬於東海系群，而東海系群的行動模式介於洄游與非洄游之間。2000年為止，在土佐灣雖已辨識出43隻個體，而聲音和個體辨識上的研究能幫助我們更了解土佐灣的族群。本研究在2008年8~12月記錄並分析了兩種聲音類型，分別為可能與攝食行為有關的聲音類型Be8a，以及可能為母子分離時維持聯繫相關的nonpulsed moan。另外在此研究期間，除了2008年的布氏鯨照片外，同時蒐集2001~2007年的照片供個體辨識，共有45隻個體被辨識出，其中有14隻個體是2000年之前已被辨識之個體。土佐灣終年可目擊布氏鯨，但春夏多在土佐灣中部目擊並有攝食行為，而秋冬則多在西南方目擊並有求偶行為。本研究認為東海系群的布氏鯨可能分為兩個(或以上)社群，而土佐灣的布氏鯨屬於北方社群。

Bryde’s whale (Balaenoptera edeni) is a species of baleen whales widely distributed throughout the tropical and temperate waters and does not migrate for long distance. Phylogenetically, the population in Tosa Bay, which includes about 43 individuals identified, belongs to the East China Sea Stock and it behaves between migratory and non-migratory forms. Studies on acoustic signals and individual identification could help us understand Bryde’s whales in Tosa Bay more. Between August and December 2008, two sound types were recorded and analyzed. The sound type Be8a was considered as feeding call, while the nonpulsed moan was considered associated with mother-calf communication during separation. In this study period, other 31 individuals were identified and added into the catalogue. Bryde’s whales are distributed very limited. Concentration of individuals was found in central Tosa Bay between spring and early autumn and it shifted southwestern in late autumn and winter. This study suggests that the East China Sea Stock may separate into two or more communities and the population in Tosa Bay belongs to the northern community.

Acknowledgments i
Abstract ii
中文摘要 iii
Table of Contents iv
List of Tables v
List of Figures vi
I. Introduction 1
A. Bryde’s whale (Balaenoptera edeni) 1
B. Bioacoustic Studies 4
C. Individual Identification (Photo-identification) 7
D. Bryde’s whales in Tosa Bay 8
II. Purposes of This Study 9
III. Materials and Methods 10
A. Study Site 10
B. Field Survey 10
C. Recording Equipments 11
D. Acoustic Analysis 12
E. Photographic Equipments 12
F. Individual Identification 12
G. Marine Environment 13
IV. Results 15
A. Survey Effort and Sighting 15
B. Acoustic Analysis 15
C. Individual Identification 17
D. Marine Environment 20
V. Discussion 22
A. Acoustic Analysis 22
B. Individual Identification 24
C. Marine Environment 26
VI. Conclusion 28
References 29
Tables 36
Figures 40
Appendix I 60
Appendix II: Catalogue of Individual Identification for Bryde’s whales in Tosa Bay
61

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