魟、鰩有成長緩慢、性成熟晚、產仔數或產卵數少等特性，只要漁獲壓力稍大，其族群即有可能崩潰甚至滅絕之虞。由於魟、鰩管理之迫切性，卻無法對每種魟、鰩進行詳細的資源評估，因此本研究蒐集35 種41 系群之生活史特性進行分析，包含出生體長與極限體長之比值（Lb/L∞）、性成熟體長與極限體長之比值（Lm/L∞）、性成熟年齡（Tm）、成長係數（k）、最大年齡（Tmax）及年胎仔數（f/Rc）。以主成份分析得到主成份得點，再利用集群分析法進行分群，最後利用迴歸分析找出生活史參數與族群成長率之關係，並建立估計各群族群成長率（λ）之經驗式。結果顯示41 系群可分為3 群，第一群為較早熟（Lm/L∞＜0.62）、成長緩慢（k＜0.090 yr-1）、壽命長（Tmax＞28 yr），如金魟（Dasyatis chrysonota）等，估計λ 的經驗式為λ'=1.7127+0.3127*ln(Lm/L∞)-0.2720*ln(Tm)+0.1080*ln(f/Rc)；第二群為較晚熟（Lm/L∞＞0.62）、成長速率介於中間（0.090 yr-1＜k＜0.160 yr-1）、最大年齡也介於中間（17 yr＜Tmax＜28 yr），如滑鰩（Dipturus laevis）等，λ'=1.2523-0.4038*ln(Tm)-0.2302*ln(k)+0.1166*ln(f/Rc)；第三群為出生體長與極限體長之比值大（Lb/L∞＞0.18）、成長快速（k＞0.160 yr-1）、壽命短（Tmax＜17 yr），如紫魟（Pteroplatytrygon violacea）等，λ'=0.9205+0.0603*ln(Lb/L∞)-0.3744*ln(Lm/L∞)+0.1234*ln(f/Rc)。依照個別魟、鰩之生活史特性帶入本研究所發展所屬群組之迴歸方程式，即可預測該魚種之族群成長率，且與傳統人口學分析法所得之結果相近。此方法可減少因生活史參數估計之不確定性所造成族群成長率之誤差，是一經濟有效的資源管理工具。

Skates and rays generally grow slowly, reach sexual maturity at a late age, and have low fecundity. These stocks may be depleted or collapsed under heavy fishing pressure. Thus, management of these species is urgently needed. This study used the analysis of life history parameters to achieve this goal. Life history parameters including the ratio between size at birth and asymptotic length (Lb/L∞), the ratio between size at maturity and asymptotic length (Lm/L∞), maximum age (Tmax), age at maturity (Tm), growth rate (k) and the ratio between fecundity and reproduction cycle (f/Rc) of 35 species (41 stocks) were analyzed with principal component analysis (PCA) to calculate factor coordinates. Three groups were categorized by cluster analysis based on PC scores. Three empirical equations describing the relationships between population growth rates (λ' ) and life history parameters were developed as followings: (1) λ'=1.7127+0.3127*ln(Lm/L∞)-0.2720*ln(Tm)+0.1080*ln(f/Rc), for species with slow growth rate (k＜0.090 yr-1), high longevity (Tmax＞28 yr) and early maturity (Lm/L∞＜0.62), (e.g. Dasyatis chrysonota etc.); (2) λ'=1.2523-0.4038*ln(Tm)-0.2302*ln(k)+0.1166*ln(f/Rc), for species with intermediate growth rate (0.090 yr-1＜k＜0.160 yr-1), intermediate longevity (17 yr＜Tmax＜28 yr), late mature (Lm/L∞＞0.62), (e.g. Dipturus laevis etc.); (3) λ'=0.9205+0.0603*ln(Lb/L∞)-0.3744*ln(Lm/L∞)+0.1234*ln(f/Rc), for species with fast growth rate (k＞0.160 yr-1), low longevity (Tmax＜17 yr) and big size at birth (Lb/L∞＞0.18), (e.g. Pteroplatytrygon violacea etc.). The population growth rates predicted by empirical equations developed in this study have good agreement with those from conventional demographic analysis. Our empirical equations can reduce the uncertainties from life history parameter estimation and increase the accuracy of estimation of population growth rates.

第一章 前言................................................1
1-1分類地位及型態特徵........................................1
1-2緣由與動機...............................................2
1-3文獻回顧.................................................4
1-4研究目的.................................................8
第二章 材料與方法..........................................10
2-1資料來源................................................10
2-1-1生活史參數之蒐集.......................................10
2-1-2生態參數之蒐集........................................13
2-2資料處理與分析..........................................14
2-2-1自然死亡係數之估計.....................................14
2-2-2以人口學方法估計族群成長率..............................14
2-2-3輸入之生活史參數.......................................15
2-3多變量分析法............................................16
2-3-1主成份分析法..........................................16
2-3-2集群分析法............................................18
2-3-3迴歸分析法............................................18
2-4生態參數與分群..........................................19
第三章 結果...............................................20
3-1生活史參數..............................................20
3-1-1生殖參數..............................................20
3-1-2年齡成長參數..........................................21
3-1-3各體長間之比值........................................22
3-2參數之估計..............................................23
3-2-1生殖週期與年胎仔數（或年產卵數）.........................23
3-2-2最大年齡之估計........................................23
3-2-3自然死亡係數之估計.....................................24
3-2-4以人口學估計之族群成長率................................24
3-3多變量分析法............................................25
3-3-1主成份分析............................................25
3-3-1-1生活史參數之相關性...................................25
3-3-1-2特徵值與解釋變異.....................................26
3-3-1-3特徵向量............................................26
3-3-1-4生活史參數與主成份之相關性............................27
3-3-1-5主成份得點..........................................27
3-3-2集群分析..............................................28
3-3-3生態參數與分群........................................29
3-3-4迴歸分析..............................................29
3-3-4-1各群之經驗式........................................29
3-3-4-2以迴歸分析估計族群成長率..............................30
第四章 討論...............................................31
4-1生活史參數之引用.........................................31
4-1-1生殖週期與年胎仔數（或年產卵數）.........................31
4-1-2成長係數..............................................32
4-1-3體長.................................................33
4-1-4出生體長與極限體長之比值................................33
4-1-5成熟體長與極限體長之比值................................34
4-2最大年齡之估計..........................................35
4-3自然死亡係數之估計.......................................35
4-4以人口學方法估計之族群成長率..............................36
4-5輸入之生活史參數.........................................37
4-6生活史參數之相關性.......................................38
4-7多變量分析..............................................39
4-8以迴歸分析估計之族群成長率................................39
4-9生活史參數之不確定性.....................................40
4-9-1生殖參數..............................................40
4-9-2年齡成長參數..........................................41
4-10生活史特性之比較........................................42
4-11生態參數與生活史參數之關聯性.............................42
4-12資源管理之建議.........................................43
4-13結論..................................................46
4-14後續研究...............................................46
參考文獻...................................................48
附圖......................................................65
附表......................................................90

李姵萱 (2008)。台灣東北部海域耳棘老板鯆之年齡與成長。國立台灣海洋大學碩士論文。89pp。
林世華 (2002)。以人口學分析法進行台灣東北部海域深海狐鮫之資源評估。國立台灣海洋大學碩士論文。75pp。
張淑如 (2006)。台灣東北部海域耳棘老板鯆之生殖生物學研究及胃內容物分析。國立台灣海洋大學碩士論文。86pp。
張瑞涵 (2006)。西北太平洋海域灰鯖鮫之資源評估。國立台灣海洋大學碩士論文。122pp。
陳純慧 (2007)。以生活史分析進行魟、鰩資源管理之研究。國立台灣海洋大學碩士論文。131pp。
陳順宇 (2000)。迴歸分析。華泰書局。520pp。
陳順宇 (2005)。多變量分析。華泰書局。710pp。
蔡文沛 (2004)。台灣東北部海域淺海狐鮫之資源評估。國立台灣海洋大學碩士論文。111pp。
Abdel-Aziz, S. H., A. N. Khalil, and S. A. Abdel-Maguid. (1993). Reproductive cycle of the common guitarfish, Rhinobatos rhinobatos (Linnaeus, 1758), in Alexandria Waters, Mediterranean Sea. Australian Journal of Marine Freshwater Research. 40, 507-517.
Au, D. W. and S. E. Smith. (1997). A demographic method with population density compensation for estimating productivity and yield per recruit of the leopard shark (Triakis semifasciata). Canadian Journal of Fisheries and Aquatic Sciences. 54, 415-420.
Babel, J. S. (1967). Reproduction, life history, and ecology of the round stingray, Urolophus halleri Cooper. Fishery Bulletin. 137, 1-104.
Basusta, N., S. A. Demirhan, E. Cicek, A. Basusta, and T. Kuleli. (2008). Age and growth of the common guitarfish, Rhinobatos rhinobatos, in Iskenderun Bay (north-eastern Mediterranean, Turkey. Journal of the Marine Biological Association of the United Kingdom. 88(4), 837-842.
Bauchot, M. L. (1987). Raies at autres batoides. In: ‘Fiches FAO d'Identification des Especs pour les Besoins de la Peche’. (Eds M. Fisher, M. Schneider and M.-L. Bauchot) pp. 847-885. Mediterranee et Mer Noire. Zone de Peche 37. Revision 1. II:. FAO, Rome.
Berestovskii, E. G. (1994). Reproductive biology of skates of the family Rajidae in the seas of the far north. Journal of Ichthyology. 34, 26-27.
Bor, P. (2002). Egg-capsules of sharks and skates. http://www.rajidae.tmfweb.nl/rogtabel.html/
Brander, K. (1981). Disappearance of Common skate Raia batis from Irish Sea. Nature. 290, 48-49.
Branstetter, S. (1987). Age and Growth Estimates for Blacktip, Carcharhinus limbatus, and Spinner, C. brevipinna, Sharks from the Northwestern Gulf of Mexico. Copeia. 1987, 964-974.
Cailliet, G. M. (1992). Demography of the central California population of the leopard shark (Triakis semifasciata). Australian Journal of Marine and Freshwater Research. 43, 183-193.
Cailliet, G. M. and Goldman, K. J. (2004). Age determination and validationIn Chondrichthyan fishes. In ‘Biology of Sharks and Their Relatives’. (Eds J. C. Carrier, J.A. Musick and M. R. Heithaus.) pp. 399-447. (CRC Press: NewYork.).
Casey, J. M. and R. A. Myers. (1998). Near extinction of a large, widely distributed fish. Science. 281, 690-692.
Christopher, M. G., S. K. Gaichas, and D. K. Kimura. (2007). Age and growth of big skate (Raja binoculata) and longnose skate (R. rhina) in the Gulf of Alaska. Enviornmental Biology of Fishes. 80, 337-349.
Clark, R. S. (1922). Skates and rays (Raiae) No.1. Egg capsules and young. Journal of the Marine Biological Association of the United Kingdom. 12, 577-643.
Coelho, R. and K. Erzini. (2002). Age and growth of the undulate ray, Raja undulata, in the Algarve (southern Portugal). Journal of the Marine Biological Association of the United Kingdom. 82, 987-990.
Coelho, R. and K. Erzini. (2006). Reproductive aspects of the undulate ray, Raja undulata, from the south coast of Portugal. Fisheries Research. 81, 80-85.
Compagno, L. J. V. (1984). Sharks of the world. An annotated and illustrated catalogue of shark species known to data. FAO Species Catalogue. Vol. 4, Parts 1 and 2. FAO Fish. Synopsis 125. FAO, Rome, Italy.
Compagno, L. J. V. (1990). Alternative life-history styles of cartilaginous fishes in time and space. Enviornmental Biology of Fishes. 28, 33-75.
Compagno, L. J. V. (1999). Checklist of living elasmobranches. In ‘Sharks, skates, and rays: The biology of elasmobranch fishes’. (Eds W. C. Hamlett). p471-498. The John Hopkins University Press, Baltimore.
Cortes, E. (1995). Demographic analysis of the Atlantic sharpnose shark, Rhizoprionodon terraenovae, in the Gulf of Mexico. Fishery Bulletin. 93, 57-66.
Cortes, E. (1998). Demographic analysis as an aid in shark stock assessment and management. Fisheries Research. 39, 199-208.
Cortes, E. (1999). Standardized diet compositions and trophic levels of sharks. ICES Journal of Marine Science. 56, 707-717.
Cortes, E. (2000). Life-history patterns and correlations in sharks. Reviews in Fisheries Science (8), 299-344.
Cortes, E. and G. R. Parsons. (1996). Comparative demography of two populations of the bonnethead shark (Sphyrna tiburo). Canadian Journal of Fisheries and Aquatic Sciences. 53, 709-718.
Cowley, P. D. (1997). Age and growth of the blue stingray Dasyatis chrysonota chrysonota from the south-eastern cape coast of SouthAfrica. South African Journal of Marine Science. 18, 31-38.
Cox, D. L. and T. J. Koob. (1993). Predation on elasmobranch eggs. Environmental Biology of Fishes. 38, 117-125.
Davis, C. D., G. M. Cailliet, and D. A. Ebert. (2007). Age and growth of the roughtail skate Bathyraja trachura (Gilbert 1892) from the eastern North pacific. Environmental Biology of Fishes. 80, 325-336.
Del Rio, J. L. (2002). Some Aspects of the Thorny Skate, Amblryraja radiata, reproductive Biology. In NAFO Division 3N. NAFO SCR Doc. 02/118, serial no. N4739, 14p.".
Du buit, M. H. (1977). Age et croissance de Raja batis et de Raja naevus en Mer Celtique. ICES Journal of Marine Science. 37, 261-265.
Dulvy, N. K. and J. D. Reynolds. (2002). Predicting Extinction Vulnerability in Skates. Conservation Biology. 16, 440-450.
Dulvy, N. K., J. D. Reynolds, J. D. Metcalfe, and J. Glanville. (2000). Fisheries stability, local extinctions and shifts in community structure in skates. Conservation Biology. 14, 1-11.
Ebert, D. A. (2003). Sharks, Rays and Chimaeras of California. University of California Press, Berkeley, California, p. 284.
Ebert, D. A. (2005). Reproductive biology of skates, Bathyraja (Ishiyama), along the eastern Bering Sea continental slope. Journal of Fish Biology. 66, 618-649.
Ebert, D. A. and C. D. Davis. (2007). Descriptions of skate egg cases (Chondrichthyes: Rajiformes: Rajoidei) from the eastern North Pacific. Zootaxa. 1393, 1-18.
Ebert, D. A. and J. J. Bizzarro. (2007). Standardized diet compositions and trophic levels of skates (Chondrichthyes: Rajiformes: Rajoidei). Environmental Biology of Fishes. 80, 221-237.
Ebert, D. A. and L. J. V. Compagno. (2007). Biodiversity and systematics of skates (Chondrichthyes: Rajiformes: Rajoidei). Environmental Biology of Fishes. 80, 111-124.
Ebert, D. A. and P. D. Cowley. (2008). Reproduction and embryonic development of the blue stingray, Dasyatis chrysonota, in southern African waters. Journal of the Marine Biological Association of the United Kingdom.p.1-7.
Ebert, D.A., W. D. Smith, D. L. Haas, S. M. Ainsley, and G. M. Cailliet. (2007).Life history and population dynamics of Alaskan skates: providing essential biological information for effective management of bycatch and target species. North Pacific Research Board Final Report 510, 124 p.
Edwards, R. R. C. (1980). Aspect of the Population Dynamics and Ecology of the White Spotted Stingaree, Urolophus paucimaculatus Dixon, in Port Phillip Bay, Victoria. Australian Journal of Marine Freshwater Research. 31, 459-467.
Enever, R., T. L. Catchpole, J. R. Ellis, and A. Grant. (2009). The survival of skates (Rajidae) caught by demersal trawlers fishing in UK waters. Fisheries Research. 97, 72-76.
Fabens, A. J. (1965). Properties and fitting of the von Bertalanffy growth curve. Growth. 29, 265-289.
Frisk, M. G. (unpublished data). Comments on fecundity and reproductive biology of winter skate, Raja ocellata, in the western Atlantic.
Frisk, M. G. and T. J. Miller. (2005). Life Histories and Vulnerability to Exploitation of Elasmobranchs: Inferences from Elasticity, Perturbation and Phylogenetic Analyses. Journal of Northwest Atlantic Fishery Science. 35, 27-45.
Frisk, M. G., T. J. Miller, and M. J. Fogarty. (2001). Estimation and analysis of biological parameters in elasmobranch fishes: a comparative life history study. Canadian Journal of Fisheries and Aquatic Sciences. 56, 578-773.
Frisk, M. G., T. J. Miller, and M. J. Fogarty. (2002). The population dynamics of little skate Leucoraja erinacea, winter skate Leucoraja ocellata, and barndoor skate Dipturus laevis: predicting exploitation limits using matrix analyses. ICES Journal of Marine Science. 59, 576-586.
Fuentealba, M. and M. Leible. (1990). Perspectivas de la pesquer?’a de la raya volant?’n Raja (Dipturus) flavirostris: Estudio de edad, crecimiento y algunos aspectos reproductivos. In ‘Perspectivas de la Actividad Pesquera en Chile’. (Eds, M. A. Barbieri) pp. 227-236. Valparaiso: Escuela de Ciencias del Mar, Universidad Cato’ lica de Valpara?’so.Gallucci, F.V., I. G. Taylor, and K. Erzini. (2006). Conservation and management of exploited shark populations based on reproductive value. Canadian Journal of Fisheries and Aquatic Sciences 63, 931-942.
Gallucci, F.V., I. G. Taylor, and K. Erzini. (2006). Conservation and management of exploited shark populations based on reproductive value. Canadian Journal of Fisheries and Aquatic Sciences. 63, 931-942.
Garcia, V. B., L. O. Lucifora, and R. A. Myers. (���~�~��). The importance of habitat and life history to extinction risk in sharks, skates, rays and chimaeras. Proceedings of the Royal Society B: Biological Sciences. 275, 83-89.
Gedamke, T., W. D. DuPaul, and J. A. Musick. (2005). Observations on the Life History of the Barndoor Skate, Dipturus laevis, on Georges Bank (Western North Atlantic). Journal of Northwest Atlantic Fishery Science. 35, 67-78.
Gertseva, V. V. (2009). The population dynamics of the longnose skate, Raja rhina, in the northeast Pacific Ocean. Fisheries Research. 95, 146-153.
Gunderson, D. R. (1980). Using r-K selection theory to predict natural mortality. Canadian Journal of Fisheries and Aquatic Sciences. 37, 2266-2271.
Gunderson, D. R. and P. H. Dygert. (1988). Reproductive effort as a predictor of natural mortality rate. Journal du Conseil International pour L’exploration de la Mer. 44, 200-209.
Hale, L. F. and C. G. Lowe. (2008). Age and growth of the round stingray Urobatis halleri at Seal Beach, California. Journal of Fish Biology. 73, 510-523.
Hemida, F., R. Seridji, S. Ennajar, Bradai, M. N., E. Collier, O. Guelorget, and C. Capape. (2003). New observations on the reproductive biology of the pelagic stingray, Dasyatis violacea Bonaparte, 1832 (Chondrichthyes: Dasyatidae) from the Mediterranean Sea. Acta adriatica. 44(2), 193-204.
Henderson, A. C., A. I. Arkhipkin, and J. N. Chtcherbich. (2004). Distribution, Growth and Reproduction of the White-spotted Skate Bathyraja albomaculata (Norman, 1937) Around the Falkland Islands. Journal of Northwest Atlantic Fishery Science. 35, 79-87.
Hoenig, J. M. (1983). Empirical use of longevity data to estimate mortality rates. Fishery Bulletin. 82, 898-903.
Hoenig, J. M. and S. H. Gruber (1990). Life-history patterns in the elasmobranchs: implications for fisheries management. In ‘Elasmobranchs as Living Resources: Advances in the Biology, Ecology, Systematics, and the Status of the Fisheries’. (Eds H. L. Pratt, S. H. Gruber, and T. Taniuchi.) pp. 1-16. Proceedings of the Second United States-Japan Workshop East-West Center, Honolulu, Hawaii, 9-14 December 1987, NOAA Technical Report NMFS 90. 518 pp.
Hoff, T. B. (1990). Conservation and management of the western North Atlantic shark resource based on the life history strategy limitations of sandbar sharks. Ph.D dissertation. University of Delaware, Newark, DE, 282pp.
Hoff, T. B. and J. Musick. (1990). Western North Atlantic shark-fishery management problems and information requirement. In ‘Elasmobranchs as Living Resources: Advances in the Biology, Eocology, Systematics, and the Status of Fisheries’. (Eds H. L. Pratt, S. h. Gruber and T. Taniuchi) NOAA Technical Report NMFS No. 90, 455-472.
Holden, M. J. (1972). The growth rates of Raja brachyura, R. clavata and R. montagui as determined from tagging data. Journal du Conseil International pour L’exploration de la Mer. 34(2), 161-168.
Holden, M. J. (1974). Problems in the rational exploitation of elasmobranch populations and some suggested solutions. In sea fisheries research (Eds F. R, harden Jones)pp. 117-137. John Wiley and Sons, New York, NewaYork, USA.
Holden, M. J. (1975). The fecundity of Raja clavata in British waters. Journal du Conseil International pour L’exploration de la Mer. 36(2), 110-118.
Ismen, A. (2003). Age, growth, reproduction and food of common stingray (Dasyatis pastinaca L.,1785) in Iskenderun Bay, the eastern Mediterranean. Fisheries Research. 60, 169-176.
Ismen, A., C. Yigin and P. Ismen. (2007). Age, growth, reproductive biology and feed of the common guitarfish (Rhinobatos rhinobatos Linnaeus, 1758) in Iskenderun Bay, the eastern Mediterranean Sea. Fisheries Research. 84, 263-269.
Izzo, C. and B. M. Gillanders. (2008). Initial assessment of age, growth and reproductive parameters of the southern fiddler ray Trygonorrhina fasciata (Muller & Henle, 1841) from South Australia. Pan-American Journal of Aquatic Sciences. 3(3), 321-327.
Jennings, S., S. P. R. Greenstreet, and J. D. Reynolds. (1999). Structural change in an exploited fish community: aconsequence of differential fishing effects on species with contrasting life histories. Journal of Animal Ecology (68), 617-627.
Johnson, G. F. (1979). The biology of the little skate, Raja erinacea Michill 1825, in Block Island Sound, Rhode Island. Thesis: University of Rhode Island, 118 pp.
Joung, S. J. (1993). Biology of the sandbar shark, Carcharhinus plumbeus,in northeastern waters of Taiwan. Ph. D. Dissertation, National Taiwan Ocean University Keelung, Taiwan. 162pp.
King, J. R and G. A. Mcfarlane. (2003). Marine fish life history strategies: applications to fishery management. Fisheries Management and Ecology. 10, 249-264.
Krebs, C. J. (1985). Ecology: the experimental analysis of distribution and abundance, 3rded. Harper and Row, New York, NY, 800pp.
Laptikhovsky, V. V. (2004). Survival rates for rays discarded by the bottom trawl squid fishery off the Falkland Islands. Fishery Bulletin. 102, 757-759.
Licandeo, R. R., J. G. Lamilla, P. G. Rubilar, and R. A. Vega. (2006). Age, growth, and sexual maturity of the yellownose skate (Dipturus chilensis) off the southeastern Pacific. Journal of Fish Biology. 68, 488-506.
Licandeo, R., F. Cerna, and R. Cespedes. (2007). Age, growth, and reproduction of the roughskin skate, Dipturus trachyderma, from the southeastern Pacific. ICES Journal of Marine Science. 64, 141-148.
Liu, K. M. and C. T. Chen. (1999). Demographic analysis of the scalloped hammerhead, Sphyrna lewini, in the northwestern Pacific. Fisheries Science. 65, 218-223.
Lucifora, L. O. and V. B. Garcia. (2004). Gastropod predation on egg cases of skates (Chondrichthyes, Rajidae) in the southwestern Atlantic: quantification and life history implications. Marine Biology. 145, 917-922.
Mariano-Melendez, E. (1997). Biologia reproductiva de la raya lodera Dasyatis brevis (Garman, 1880), en Bahia Almejas, B.C.S., Mexico. Tesis, Universidad Automa de Baja California Sur, La Paz, B.C.S., Mexico.
Marshall, L. J., W. T. White, and I. C. Potter. (2007). Reproductive biology and diet of the southern fiddler ray, Trygonorrhina fasciata (Batoidea Rhinobatidae) an important trawl bycatch species. Marine and Freshwater Research. 58, 104-115.
Martin, L. K. and G. M. Cailliet. (1988a). Aspects of the reproduction of the Bat ray, Myliobatis californica, in central california. Copeia. 1988(3), 754-762.
Martin, L. K. and G. M. Cailliet. (1988b). Age and growth determination of the Bat Ray, Myliobatis californica Gill, in central california. Copeia. 1988(3): 762-773.
Matta, E. M. and D. R. Gunderson (2007). Age, growth, maturity, and mortality of the Alaska skate, Bathyraja parmifera, in the eastern Bering Sea. Environmental Biology of Fishes. 80, 309-323.
McEachran, J. D. (2002). Skates. Family Rajidae. In ‘Bigelow and Schroeder's fishes of the Gulf of Maine’. (Eds B. B. Collette and G. Klein-MacPhee)3rd Edition. pp. 60-75. Smithsonian Institution Press, Washington, DC. 748 pp.
McFarlane, G. A. and J. R. King. (2006). Age and growth of big skate (Raja binoculata) and longnose skate (Raja rhina) in British Columbia waters. Fisheries Research. 78, 169-178.
Mollet, H. F. and G. M. Cailliet. (2002). Comparative population demography of elasmobranches using life history tables, Leslie matrices and stage-based matrix models. Marine and Freshwater Research. 53, 503-516.
Mollet, H. F., J. M. Ezcurra, and J. B. O'Sullivan. (2002). Captive biology of the pelagic stingray, Dasyatis violacea (Bonaparte, 1832). Marine and Freshwater Research. 53, 531-541.
Musick, J. A. (1999). Ecology and conservation of long-lived marine animals. In ‘American Fisheries Society Symposium 23: Life in the Slow Lane: Ecology and Conservation of Long-lived Marine Animals’. (Ed. J. A. Musick.) pp. 1-10. (American Fisheries Society: Bethesda, MD.)
Musick, J.A. and J. K. Ellis. (2005). Reproductive evolution of chondrichthyans. In ‘Reproductive biology and phylogeny of Chondrichthyes: sharks, batoids, and chimaeras’. (Eds W.C. Hamlett) pp. 45-79. Enfield, NH: Science Publishers,.
Muus, B. J. and J. G. Nielsen. (1999). Sea fish. In ‘Scandinavian Fishing Year Book’, Hedehusene, Denmark, 340p.
Neer, J. A. and B. A. Thompson. (2005). Life history of the cownose ray, Rhinoptera bonasus, in the northern Gulf of Mexico, with comments on geographic variability in life history traits. Environmental Biology of Fishes. 73, 321-331.
Neer, J. A. and G. M. Cailliet. (2001). Aspects of the life history of the Pacific electric ray, Torpedo californica (Ayers). Copeia. 2001, 842-847.
Neer, J. A., B. A. Thompson and J. K. Carlson. (2005). Age and growth of Carcharhinus leucas in the northern Gulf of Mexico: incorporating variability in size at birth. Journal of Fish Biology. 67: 370-383.
Nelson, J. S. (2006). ‘Fishes of the world.’ John Wiley and Sons, Inc. New York. 4th edition. 601 pp.
Pauly, D. (1980). On the interrelationships between natural mortality, growth parameters, and mean environmental temperature in 175 fish stocks. Journal du Conseil International pour L’exploration de la Mer. 39, 175-192.
Pratt, H. L. Jr. and J. G. Casey (1990). Shark reproductive strategies as a limiting factor in directed fisheries, with a review of Holden's method of estimating growth-parameters. NOAA Technical Report, NMFS 90: 98-129.
Quiroz, J. C. and R. Wiff. (2005). Demographic analysis and exploitation vulnerability of beaked skate (Dipturus chilensis) off the Chilean austral zone. ICES CM 2005/N:19
Revill, A. S., N. K. Dulvy, and R. Holst. (2005). The survival of discarded lesser-spotted dogfish (Scyliorhinus canicula) in the Western English Channel beam trawl fishery. Fisheries Research. 71, 121-124.
Richards, S. W., D. Merriman, and L. H. Calhoun. (1963). Studies on the marine resources of southern New England, IX. The biology of the little skate, Raja erinacea Mitchill. Bulletin of the Bingham Oceanographic Collection, Yale University. 18(3):5-67.
Ricker, D. A. (1979). Growth rates and Models. In ‘Fish physiology, Volume VIII’. (Eds W. S. Hoar and D. J. Randall) pp. 677-743. (Academic Press: New York.)
Rijnsdorp, A. D., N. Daan, W. Dekker, J. J. Poos, and W. L. T. Van Densen. (2007). Sustainable use of flatfish resources: Addressing the credibility crisis in mixed fisheries management. Journal of Sea Research. 57, 114-125.
Ryland, J. S. and T. O. Ajyi. (1984). Growth and population dynamics of three Raja species (Batoidei) in Carmarthen Bay, British Isles. Journal du Conseil International pour L’exploration de la Mer. 41, 111-120.
Schindler, D. E., T. E. Essington, J. F. Kitchell, C. Boggs, and R. Hilborn. (2002). Sharks and tunas: fisheries impacts on predators with contrasting life histories. Ecological Applications. 12, 735-748.
Serena, F. (2005). Field identification guide to the sharks and rays of the Mediterranean and Black Sea. FAO Species Identification Guide For Fishery Purposes. Rome: FAO.
Serena, F., M. Barone, C. Mancusi, and A. J. Abella. (2005). Reproductive biology, growth and feeding habits of Raja Asterias delaroche, 1809, from the north tyrrhenian and south ligurian sea (Italy), with some notes on trends in landings. 2005 ICES Annual Science Conference; Theme Session on Elasmobranch Fisheries Science: 20-24 September 2005. CM2005/N:20
Simpfendorfer, C. A. (1999). Mortality estimates and demographic analysis for the Australian sharpnose shark, Rhizoprionodon taylori, from northern Australia. Fishery Bulletin. 97, 978-986.
Simpfendorfer, C. A. (2000). Predicting population recovery rates for endangered western Atlantic sawfishes using demographic analysis. Environmental Biology of Fishes. 58, 371-377.
Sminkey, T. R. and J. A. Musick. (1996). Age and growth of the sandbar shark, Carcharhinus plumbeus, before and after population depletion. Fishery Bulletin. 94, 871-883.
Smith, J. W. and J. V. Merriner. (1986). Observations on the reproductive biology of the Cownose Ray, Rhinoptera bonasus, in Chesapeake Bay. Fishery Bulletin. 84, 871-877.
Smith, J. W. and J. V. Merriner. (1987). Age and Growth, Movements and Distribution of the Cownose Ray, Rhinoptera bonasus, in Chesapeake Bay. Estuarine Research Federation. 2, 153-164.
Smith, W. D. (2005). Life history aspects and population dynamics of a commercially exploited stingray, Dasyatis dipterura. M.Sc. Thesis, Moss Landing Marine Laboratories, California, USA.
Smith, W. D., G. M. Cailliet, and E. Cortes (2008). Demography and elasticity of the diamond stingray, Dasyatis dipterura: parameter uncertainty. Marine and Freshwater Research. 59, 575-586.
Smith, W. D., G. M. Cailliet, and E. M. Melendez. (2007). Maturity and growth characteristics of a commercially exploited stingray, Dasyatis dipterura. Marine and Freshwater Research. 58, 54-66.
Stevens, J. D., R. Bonfil, N. K. Dulvy, and P. A. Walker. (2000). The effects of fishing on sharks, rays, and chimaeras (chondrichthyans), and the implications for marine ecosystems. ICES Journal of Marine Science. 57, 476–494.
Sulikowski, J. A., J. Kneebone, and S. Elzey. (2005a). Age and growth estimates of the thoeny skate (Amblyraja radiata) in the western Gulf of Maine. Fishery Bulletin. 103, 161-168.
Sulikowski, J. A., J. Kneebone, and S. Elzey. (2005b). The reproductive cycle of the thorny skate (Amblyraja radiata) in the western Gulf of Maine. Fishery Bulletin. 103, 536-543.
Sulikowski, J. A., J. Kneebone, S. Elzey, J. Jurek, W. H. Howell, and P. C. W. Tsang. (2006). Using the composite variables of reproductive morphology, histology and steroid hormones to determine age and size at sexual maturity for the thorny skate Amblyraja radiata in the western Gulf of Maine. Journal of Fish Biology. 69, 1449-1465.
Sulikowski, J. A., M. D. Morin, S. H. Suk, and W. H. Howell. (2003). Age and growth estimate of the winter skate (Leuoraja ocellata) in the western Gulf of Maine. Fishery Bulletin. 101, 405-413.
Sulikowski, J. A., P. C. W. Tsang, and W. H. Howell. (2005c). Age and size at sexual maturity for the winter skate, Leucoraja ocellata, in the western Gulf of Maine based on morphological, histological and steroid hormone analyses. Environmental Biology of Fishes. 72, 429-441.
Takeuchi, Y., Y. Senba, and H. Nakano. (2005). Demographic analysis on Atlantic blue and shortfin mako sharks. Collective Volume of Scientific Papers ICCAT. 58, 1157-1165.
Taylor, C. C. (1958). Cod growth and temperature. Journal du Conseil International pour L’exploration de la Mer. 23, 366-370.
Thorburn, Dean C. and D. L. Morgan. (2005). Threatened fishes of the world: Pristis microdon Latham 1794 (Pristidae). Environmental Biology of Fishes. 72, 465-466.
Thorburn, Dean C., D. L. Morgan, A. J. Rowland, H. S. Gill, and E. Paling. (2007). Life history notes of the critically endangered dwarf sawfish, Pristis clavata, Garman 1906 from the Kimberley region of Western Australia. Enviornmental Biology of Fishes. 83, 139-145.
Thorson, J. T. and C. A. Simpfendorfer. (2009). Gear selectivity, and sample size effects on growth curve selection in shark age and growth studies, Fisheries Research. 98, 75-84.
Trippel, E. A., O. S. Kjebu, and P. Solemial. (1997). Effects of adult age and size structure on reproductive output in marine fishes. In: ‘Early life history and Recruitment in Fish Population.’ (Eds R. C. Chambers, E. A. Trippel) Chapmen and Hall, London. p.26-62.
Walker, P. A. and J. R. G. Hislop. (1998). Sensitive skates or resilient rays? Spatial and temporal shifts in ray species composition in the central and north-western North Sea between 1930 and the present day. ICES Journal of Marine Science. 55, 392-402.
Waring, G. T. (1984). Age, growth and mortality of the little skate off the northeast coast of the United States. Transactions of the American Fisheries Society. 113, 314-321.
White, W. T. and I. C. Potter. (2005). Reproductive biology, size and age compositions and growth of the batoid Urolophus paucimaculatus, including comparisons with other species of the Urolophidae. Marine and Freshwater Research. 56, 101-110.
White, W. T., M. E. Hall, and I. C. Potter. (2001). Relationship between reproductive biology and age composition and growth in Urolophus lobatus (Batoidea: Urolophidae). Marine Biology. 138, 135-147.
White, W. T., N. G. Hall, and I. C. Potter. (2002). Reproductive biology and growth during pre- and postnatal life of Trygonoptera mucosa and Trygonoptera personata (Batoidea: Urolophidae). Marine Biology. 140, 699-712.
Whittamore, J. M. and I. D. McCarthy. (2005). The population biology of the thornback ray, Raja clavata in Caernarfon Bay, north Wales. Journal of the Marine Biological Association of the United Kingdom. 85, 1089-1094.
Winemiller, K. O. and K. A. Rose. (1992). Patterns of life-history diversification in North American fishes: implications for population regulation. Canadian Journal of Fisheries and Aquatic Sciences. 49(10), 2196-2218.
Wourms, J. P. (1997). Reproduction and development in chondrichthyan fishes. American Zoologist. 17, 379-410.
Yamaguchi, A. (2006). Reproductive biology of longheaded eagle ray, Aetobatus flagellum, in Ariake Sound, Kyushu, Japan. 2006 Joint Meeting of Ichthyologists and Herpetologists, July 12-17, 2006, New Orleans.
Yamaguchi, A., I. Kawahara, and S. Ito. (2005). Occurrence, growth and food of longheaded eagle ray, Aetobatus flagellum, in Ariake Sound, Kyushu, Japan. Environmental Biology of Fishes. 74(2), 229-238.